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Networks of filamentous proteins play a crucial role in cell mechanics. These cytoskeletal networks, together with various crosslinking and other associated proteins largely determine the (visco)elastic response of cells. In this letter we…

无序系统与神经网络 · 物理学 2009-11-10 D. A. Head , A. J. Levine , F. C. MacKintosh

We examine the equilibrium fluctuation spectrum of a constituent semiflexible filament segment in a network. The effect of this cross linking is to modify the mechanical boundary conditions at the end of the filament. We consider the effect…

软凝聚态物质 · 物理学 2020-01-22 Jonathan Kernes , Alex J. Levine

Disordered networks of semiflexible filaments are common support structures in biology. Familiar examples include fibrous matrices in blood clots, bacterial biofilms, and essential components of cells and tissues of plants, animals, and…

软凝聚态物质 · 物理学 2023-09-28 Saamiya Syed , Fred C. MacKintosh , Jordan L. Shivers

The kinetics of the assembly of semiflexible filaments through end-to-end annealing is key to the structure of the cytoskeleton, but is not understood. We analyze this problem through scaling theory and simulations, and uncover a regime…

软凝聚态物质 · 物理学 2023-12-18 Valerio Sorichetti , Martin Lenz

Semiflexible polymers such as filamentous actin play a vital role in the mechanical behavior of cells, yet the basic properties of cross-linked F-actin networks remain poorly understood. To address this issue, we have performed numerical…

软凝聚态物质 · 物理学 2009-11-10 D. A. Head , A. J. Levine , F. C. MacKintosh

The semiflexible F-actin network of the cytoskeleton is cross-linked by a variety of proteins including filamin, which contain Ig-domains that unfold under applied tension. We examine a simple semiflexible network model cross-linked by such…

软凝聚态物质 · 物理学 2009-11-11 B. A. DiDonna , Alex J. Levine

The cytoskeleton of eukaryotic cells provides mechanical support and governs intracellular transport. These functions rely on the complex mechanical properties of networks of semiflexible protein filaments. Recent theoretical interest has…

软凝聚态物质 · 物理学 2012-11-01 M. Atakhorrami , G. H. Koenderink , J. F. Palierne , F. C. MacKintosh , C. F. Schmidt

Myosin motor proteins drive vigorous steady-state fluctuations in the actin cytoskeleton of cells. Endogenous embedded semiflexible filaments such as microtubules, or added filaments such as single-walled carbon nanotubes are used as novel…

生物物理 · 物理学 2016-06-21 J. Gladrow , N. Fakhri , F. C. MacKintosh , C. F. Schmidt , C. P. Broedersz

We present a theoretical framework for the linear and nonlinear visco-elastic properties of reversibly crosslinked networks of semiflexible polymers. In contrast to affine models where network strain couples to the polymer end-to-end…

软凝聚态物质 · 物理学 2016-07-06 Jan Plagge , Andreas Fischer , Claus Heussinger

The cytoskeleton is an important subsystem of cells that is involved for example in cell division and locomotion. It consists of filaments that are cross-linked by molecular motors that can induce relative sliding between filaments and…

统计力学 · 物理学 2007-08-08 P. K. Mohanty , K. Kruse

The actin cytoskeleton in living cells has many types of crosslinkers. The mechanical interplay between these different crosslinker types is an open issue in cytoskeletal mechanics. We develop a framework to study the cooperativity and…

软凝聚态物质 · 物理学 2015-05-28 Moumita Das , D. A. Quint , J. M. Schwarz

Here, we provide an overview of theoretical approaches to semiflexible polymers and their networks. Such semiflexible polymers have large bending rigidities that can compete with the entropic tendency of a chain to crumple up into a random…

软凝聚态物质 · 物理学 2016-05-25 Chase P. Broedersz , Fred. C. MacKintosh

The phase behavior of charged rods in the presence of inter-rod linkers is studied theoretically as a model for the equilibrium behavior underlying the organization of actin filaments by linker proteins in the cytoskeleton. The presence of…

软凝聚态物质 · 物理学 2016-08-31 Itamar Borukhov , Robijn F. Bruinsma , William M. Gelbart , Andrea J. Liu

We develop a model for gels and entangled solutions of semiflexible biopolymers such as F-actin. Such networks play a crucial structural role in the cytoskeleton of cells. We show that the rheologic properties of these networks can result…

凝聚态物理 · 物理学 2009-10-28 F. C. MacKintosh , J. Kas , P. A. Janmey

We consider an explicit model of a semiflexible filament moving in two dimensions on a gliding assay of motor proteins, which attach to and detach from filament segments stochastically, with a detachment rate that depends on the local load…

软凝聚态物质 · 物理学 2019-05-23 Nisha Gupta , Abhishek Chaudhuri , Debasish Chaudhuri

Motivated by recent experiments showing nonlinear elasticity of in vitro networks of the biopolymer actin cross-linked with filamin, we present an effective medium theory of flexibly cross-linked stiff polymer networks. We model such…

软凝聚态物质 · 物理学 2008-10-19 C. P. Broedersz , C. Storm , F. C. MacKintosh

The mechanical properties of cells are dominated by the cytoskeleton, an interconnected network of long elastic filaments. The connections between the filaments are provided by crosslinking proteins, which constitute, next to the filaments,…

软凝聚态物质 · 物理学 2012-10-09 Claus Heussinger

Aiming at the mechanical properties of cross-linked biopolymers, we set up and analyze a model of two weakly bending wormlike chains subjected to a tensile force, with regularly spaced inter-chain bonds (cross-links) represented by harmonic…

生物物理 · 物理学 2014-06-24 Alice von der Heydt , Daniel Wilkin , Panayotis Benetatos , Annette Zippelius

In cells, cytoskeletal filament networks are responsible for cell movement, growth, and division. Filaments in the cytoskeleton are driven and organized by crosslinking molecular motors. In reconstituted cytoskeletal systems, motor activity…

We consider a model of an extensible semiflexible filament moving in two dimensions on a motility assay of motor proteins represented explicitly as active harmonic linkers. Their heads bind stochastically to polymer segments within a…

生物物理 · 物理学 2021-08-31 Amir Shee , Nisha Gupta , Abhishek Chaudhuri , Debasish Chaudhuri
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