基于存在数据推断海洋环境中领袖-随从行为：以珊瑚熊鲨鱼为例

Social interactions are fundamental in animal groups, including humans, and can take various forms, such as competition, cooperation, or kinship. Understanding these interactions in marine environments has been historically challenging due to data collection difficulties. However, advancements in acoustic telemetry now enable remote analysis of such behaviors. This study proposes a method to derive leader-follower networks from presence data collected by a single acoustic receiver at a specific location. Using the Kolmogorov-Smirnov distance, the method analyzes lag times between consecutive presences of individuals to infer directed relationships. Tested on simulated data, it was then applied to detection data from acoustically tagged reef manta rays (Mobula alfredi) frequenting a known site. Results revealed temporal patterns, including circadian rhythms and burst-like behavior with power-law distributed time gaps between presences. The inferred leader-follower network highlighted key behavioral patterns: females followed males more often than expected, males showed stronger but fewer associations with specific females, and smaller individuals followed others less consistently than larger ones. These findings align with ecological insights, revealing structured social interactions and providing a novel framework for studying marine animal behavior through network theory.