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We propose a model for the formation of chromatin loops based on the diffusive sliding of a DNA-bound factor which can dimerise to form a molecular slip-link. Our slip-links mimic the behaviour of cohesin-like molecules, which, along with…

生物物理 · 物理学 2017-10-04 C. A. Brackley , J. Johnson , D. Michieletto , A. N. Morozov , M. Nicodemi , P. R. Cook , D. Marenduzzo

Condensation of hundreds of mega-base-pair-long human chromosomes in a small nuclear volume is a spectacular phenomenon. This process is driven by the formation of chromosome loops. ATP consuming motor, condensin, interacts with chromatin…

软凝聚态物质 · 物理学 2021-10-27 Ryota Takaki , Atreya Dey , Guang Shi , Dave Thirumalai

We use Brownian dynamics simulations to study the formation of chromatin loops through diffusive sliding of slip-link-like proteins, mimicking the behaviour of cohesin molecules. We recently proposed that diffusive sliding is sufficient to…

软凝聚态物质 · 物理学 2020-11-02 Andrea Bonato , Chris A. Brackley , James Johnson , Davide Michieletto , Davide Marenduzzo

Chromatin loop extrusion by Structural Maintenance of Chromosome (SMC) complexes is thought to underlie intermediate-scale chromatin organization inside cells. Motivated by a number of experiments suggesting that nucleosomes may block loop…

Active loop extrusion - the process of formation of dynamically growing chromatin loops due to the motor activity of DNA-binding protein complexes - is firmly established mechanism responsible for chromatin spatial organization at different…

统计力学 · 物理学 2023-01-11 Sergey Belan , Vladimir Parfenyev

We develop a kinetic theory of cohesin-driven loop extrusion on a disordered chromatin track with transient barriers. In the stationary state, the mean loop size is shown to obey a universal law determined by the bare processivity and a…

软凝聚态物质 · 物理学 2026-04-20 A. Chervinskaya , R. Metzler , K. E. Polovnikov

Transcriptional noise, or heterogeneity, is important in cellular development and in disease. The molecular mechanisms driving it are, however, elusive and ill-understood. Here, we use computer simulations to explore the role of 3D…

Loop extrusion convincingly describes how certain Structural Maintenance of Chromosome (SMC) proteins mediate the formation of large DNA loops. Yet, most of the existing computational models cannot reconcile recent in vitro observations…

软凝聚态物质 · 物理学 2022-01-12 Andrea Bonato , Davide Michieletto

An active loop-extrusion mechanism is regarded as the main out--of--equilibrium mechanism responsible for the structuring of megabase-sized domains in chromosomes. We developed a model to study the dynamics of the chromosome fibre by…

生物大分子 · 定量生物学 2020-09-30 M. Crippa , Y. Zhan , G. Tiana

Loop extrusion by motor proteins mediates the attractive interactions in chromatin on the length scale of megabases, providing the polymer with a well-defined structure and at the same time determining its dynamics. The mean square…

生物大分子 · 定量生物学 2023-07-12 E. Marchi , Y. Zhan , G. Tiana

Motivated by the recent experimental observations of the DNA loop extrusion by protein motors, in this paper we investigate the statistical properties of the growing polymer loops within the ideal chain model. The loop conformation is…

统计力学 · 物理学 2021-04-30 Dmitry Starkov , Vladimir Parfenyev , Sergey Belan

Chromatin fibers composed of DNA and proteins fold into consecutive loops to form rod-shaped chromosome in mitosis. Although the loop growth dynamics is investigated in several studies, its detailed processes are unclear. Here, we describe…

软凝聚态物质 · 物理学 2022-05-11 Hiroshi Yokota , Masashi Tachikawa

DNA loop extrusion is arguably one of the most important players in genome organization. The precise mechanism by which loop extruding factors (LEFs) work is still unresolved and much debated. One of the major open questions in this field…

软凝聚态物质 · 物理学 2023-09-18 Andrea Bonato , Jae-Won Jang , Kyoung-Wook Moon , Davide Michieletto , Je-Kyung Ryu

Chromosomes are crumpled polymer chains further folded into a sequence of stochastic loops via loop extrusion. While extrusion has been verified experimentally, the particular means by which the extruding complexes bind DNA polymer remains…

软凝聚态物质 · 物理学 2023-06-14 Kirill Polovnikov , Bogdan Slavov

Structural Maintenance Complexes (SMC) are energy consuming motors that are important in folding the genome by loop extrusion (LE) in all stages of the cell cycle. Single molecule magnetic tweezer pulling experiments have revealed that…

软凝聚态物质 · 物理学 2026-05-07 Atreya Dey , Guang Shi , Ryota Takaki , D. Thirumalai

The spatial organization of DNA involves DNA loop extrusion and the formation of protein-DNA condensates. While the significance of each process is increasingly recognized, their interplay remains unexplored. Using molecular dynamics…

生物物理 · 物理学 2025-01-31 Ryota Takaki , Yahor Savich , Jan Brugués , Frank Jülicher

Molecular motors are involved in key transport processes inside actin-based cellular protrusions. The motors carry cargo proteins to the protrusion tip which participate in regulating the actin polymerization, and play a key role in…

生物物理 · 物理学 2014-06-12 Itai Pinkoviezky , Nir S Gov

Loop extrusion is one of the main processes shaping chromosome organisation across the cell cycle, yet its role in regulating DNA entanglement and nucleoplasm viscoelasticity remains overlooked. We simulate entangled solutions of linear…

软凝聚态物质 · 物理学 2025-02-03 Filippo Conforto , Yair Augusto Gutierrez Fosado , Davide Michieletto

Many researchers have been encouraged to describe the dynamics of chromosomal loci in chromatin using the classical Rouse model of polymer dynamics by the agreement between the measured mean square displacement (MSD) versus time of…

Cell crawling requires the generation of intracellular forces by the cytoskeleton and their transmission to an extracellular substrate through specific adhesion molecules. Crawling cells show many features of excitable systems, such as…

生物物理 · 物理学 2020-06-02 Pierre Sens
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