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Flight is a complicated task at small scales in part due to the ubiquitous unsteady air which contains it. Flying organisms deal with these difficulties using active and passive control mechanisms to steer their body motion. Body attitudes…

The natural wind environment that volant insects encounter is unsteady and highly complex, posing significant flight control and stability challenges. Unsteady airflows can range from structured chains of discrete vortices shed in the wake…

Biological Physics · Physics 2016-03-03 Sridhar Ravi , Dmitry Kolomenskiy , Thomas Engels , Kai Schneider , Chun Wang , Joern Sesterhenn , Hao Liu

Birds rely on active high-acceleration morphing and flapping to navigate complex airflows, but they can also maintain stable fixed-wing postures under persistent atmospheric disturbances. Here, we show that avian wings exhibit aerodynamic…

Fluid Dynamics · Physics 2025-07-22 Lunbing Chen , Suyang Qin , Jinpeng Huang , Yufei Yin , Yang Xiang , Hong Liu

Flapping insects are remarkably agile fliers, adapted to a highly turbulent environment. We present a series of high resolution numerical simulations of a bumblebee interacting with turbulent inflow. We consider both tethered and free…

Insects use flight muscles attached at the base of the wings to produce impressive wing flapping frequencies. The maximum power output of these flight muscles is insufficient to maintain such wing oscillations unless there is good elastic…

Fluid Dynamics · Physics 2016-07-20 Yangyang Huang , Eva Kanso

Saving energy and enhancing performance are secular preoccupations shared by both nature and human beings. In animal locomotion, flapping flyers or swimmers rely on the flexibility of their wings or body to passively increase their…

Biological Physics · Physics 2011-08-30 Sophie Ramananarivo , Ramiro Godoy-Diana , Benjamin Thiria

In windy conditions, the air is turbulent. The strong and intermittent velocity variations of turbulence are invisible to flying animals. Nevertheless, flying animals, not much larger than the smallest scales of turbulence, manage to…

Fluid Dynamics · Physics 2023-04-25 Bardia Hejazi , Christian Küchler , Gholamhossein Bagheri , Eberhard Bodenschatz

Hummingbirds and insects achieve outstanding flight performance by adapting their flapping motion to the flight requirements. Their wing kinematics can change from smooth flapping to highly dynamic waveforms, generating unsteady aerodynamic…

Fluid Dynamics · Physics 2024-08-07 Romain Poletti , Andre Calado , Lilla K. Koloszar , Joris Degroote , Miguel A. Mendez

Wing flexibility plays an essential role in the aerodynamic performance of insects due to the considerable deformation of their wings during flight under the impact of inertial and aerodynamic forces. These forces come from the complex wing…

Fluid Dynamics · Physics 2020-07-20 Hung Truong , Thomas Engels , Dmitry Kolomenskiy , Kai Schneider

Turbulence grounds aircraft and combating it in flight requires energy, yet volant wildlife fly effortlessly even on windy days. The nature of the interactions between soaring birds and transient turbulent gusts is not clear, especially…

Biological Physics · Physics 2024-12-04 Dipendra Gupta , David Brandes , Michael J Lanzone , Tricia Miller , Gregory P Bewley

This paper investigates stability analysis of flapping flight. Due to time-varying aerodynamic forces, such systems do not display fixed points of equilibrium. The problem is therefore approached via a limit cycle analysis based on Floquet…

The aerial environment in the operating domain of small-scale natural and artificial flapping wing fliers is highly complex, unsteady and generally turbulent. Considering flapping flight in an unsteady wind environment with a periodically…

Inverted landing is a challenging feat to perform in aerial robots, especially without external positioning. However, it is routinely performed by biological fliers such as bees, flies, and bats. Our previous observations of landing…

Robotics · Computer Science 2022-03-04 Bryan Habas , Bader AlAttar , Brian Davis , Jack W. Langelaan , Bo Cheng

Compared with fixed-wing flight, flapping flight can generate a higher lift and is also more maneuverable, largely resulting from the benefits of wing rotation. By analyzing the real wing kinematics of fruit flies, we found that the wing…

Fluid Dynamics · Physics 2019-06-19 Yang Xiang , Haotian Hang , Hong Liu

Existing research has yet to reach a consensus on whether and how small flying animals utilize elastic energy storage mechanisms to reduce flight energy expenditure, and there is a lack of systematic and universal methods for assessment. To…

Biological Physics · Physics 2025-10-06 Shijie Sheng , Jianghao Wu , Renxuan Bo , Long Chen , Yanlai Zhang

Aerial insects exhibit highly agile maneuvers such as sharp braking, saccades, and body flips under disturbance. In contrast, insect-scale aerial robots are limited to tracking non-aggressive trajectories with small body acceleration. This…

Robotics · Computer Science 2025-08-06 Yi-Hsuan Hsiao , Andrea Tagliabue , Owen Matteson , Suhan Kim , Tong Zhao , Jonathan P. How , YuFeng Chen

In flapping flight, motion of the wings through the air generates the majority of the force and torque that controls the body motion. On the other hand, it is not clear how much effect the body motion imposes on the wings. We investigated…

Fluid Dynamics · Physics 2015-02-25 Samane Zeyghami , Haibo Dong

The ability to fly through openings in vegetation allows insects like bees to access otherwise unreachable food sources. The specific visual strategies employed by flying insects during aperture negotiation tasks remain unknown. In this…

Biological Physics · Physics 2026-04-30 Timothy Jakobi , Matt Garratt , Mandayam Srinivasan , Sridhar Ravi

Biological studies show that hummingbirds can perform extreme aerobatic maneuvers during fast escape. Given a sudden looming visual stimulus at hover, a hummingbird initiates a fast backward translation coupled with a 180-degree yaw turn,…

Robotics · Computer Science 2019-02-27 Fan Fei , Zhan Tu , Jian Zhang , Xinyan Deng

Turbulence is omnipresent in the atmosphere and a long-standing scientific conundrum that makes flight complex. This complexity is little understood; surprisingly, when turbulence arises, air vehicles struggle while birds seem to thrive.…

Fluid Dynamics · Physics 2026-01-01 Ariane Gayout , David Lentink
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