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Wing flapping is one of the most widespread propulsion methods found in nature; however, the current understanding of the aerodynamics in bird wakes is incomplete. The role of the unsteady motion in the flow and its contribution to the…

Hovering insects are limited by their physiology and need to rotate their wings at the end of each back and forth motion to keep the wing's leading edge ahead of its trailing edge. The wing rotation at the end of each half-stroke pushes the…

Fluid Dynamics · Physics 2022-10-28 Swathi Krishna , Alexander Gehrke , Karen Mulleners

Tail bending associated with maneuvering flight of insects is a known phenomenon although there are only a few studies which analyze and quantify the effects and benefits of body configuration changes. We hypothesized that these…

Biological Physics · Physics 2017-07-26 Ayodeji T. Bode-Oke , Samane Zeyghami , Haibo Dong

Flapping insect flight is a complex and beautiful phenomenon that relies on fast, active control mechanisms to counter aerodynamic instability. To directly investigate how freely-flying D. melanogaster control their body pitch angle against…

Biological Physics · Physics 2015-03-29 Samuel C. Whitehead , Tsevi Beatus , Luca Canale , Itai Cohen

The movement of organisms is subject to a multitude of influences of widely varying character: from the bio-mechanics of the individual, over the interaction with the complex environment many animals live in, to evolutionary pressure and…

Biological Physics · Physics 2013-07-18 Friedrich Lenz , Aleksei V. Chechkin , Rainer Klages

Collective patterns of motion emerge across biological taxa: insects swarm, fish school, and birds flock. In particular, large migratory birds form strikingly ordered V-shaped formations, which experiments and direct numerical simulations…

Fluid Dynamics · Physics 2026-02-26 Olivia Pomerenk , Kenneth S. Breuer

High resolution direct numerical simulations of rotating and flapping bumblebee wings are presented and their aerodynamics is studied focusing on the role of leading edge vortices and the associated helicity production. We first study the…

Fluid Dynamics · Physics 2018-03-21 T. Engels , D. Kolomenskiy , K. Schneider , M. Farge , F. -O. Lehmann , J. Sesterhenn

The capacity for aerial maneuvering shaped the evolution of flying animals. Here we evaluate consequences of aviaian morphology for aerial performance (1,2) by quantifying static stability and control effectiveness of physical models (3)…

Populations and Evolution · Quantitative Biology 2014-10-17 Dennis Evangelista , Sharlene Cam , Tony Huynh , Austin Kwong , Homayun Mehrabani , Kyle Tse , Robert Dudley

Inspired by the wake-surfing nature of animals, this study aims to understand the aerodynamic force variation on a wing surfing in an unsteady 2-D wake. Wind tunnel experiments were conducted using Particle Image Velocimetry (PIV) and force…

Fluid Dynamics · Physics 2025-12-11 Siyang Hao , Kenneth Breuer

Unmanned underwater vehicles are increasingly employed for maintenance and surveying tasks at sea, but their operation in shallow waters is often hindered by hydrodynamic disturbances such as waves, currents, and turbulence. These unsteady…

Robotics · Computer Science 2026-02-10 Tobias Cook , Leo Micklem , Huazhi Dong , Yunjie Yang , Michael Mistry , Francesco Giorgio-Serchi

Accurate estimation of aerodynamic forces is essential for advancing the control, modeling, and design of flapping-wing aerial robots with dynamic morphing capabilities. In this paper, we investigate two distinct methodologies for force…

Robotics · Computer Science 2025-08-06 Bibek Gupta , Mintae Kim , Albert Park , Eric Sihite , Koushil Sreenath , Alireza Ramezani

The large active wing deformation is a significant way to generate high aerodynamic forces required in bat flapping flight. Besides the twisting, the elementary morphing models of a bat wing are proposed, such as wing-bending in the…

Fluid Dynamics · Physics 2015-05-15 Guan Zi-Wu , Yu Yong-Liang

Fluid dynamics, and flight in particular, is a domain where organisms challenge our understanding of its physics. Integrating the current knowledge of animal flight, we propose to revisit the use of live animals to study physical phenomena.…

Fluid Dynamics · Physics 2024-09-17 Ariane Gayout , David Lentink

This paper discusses the wing inertial effects on stability of pitch motion of hovering insects. The paper also presents a dynamic model appropriate for using averaging techniques and discusses the pitch stability results derived from the…

Classical Physics · Physics 2023-06-28 Sevak Tahmasian , Braeden Kotulak-Smith

A complete, self-contained mathematical framework for modelling the coupled aeroelastic and flight dynamic behaviour of free-flying flexible aircraft subject to atmospheric gust encounters is presented. The framework integrates three…

Computational Engineering, Finance, and Science · Computer Science 2026-04-16 Nikolaos D. Tantaroudas , Ilias Karachalios

Inspired by flight characteristics captured from live Monarch butterflies, an optimal control problem is presented while accounting the effects of low-frequency flapping and abdomen undulation. A flapping-wing aerial vehicle is modeled as…

Optimization and Control · Mathematics 2021-07-19 Tejaswi K. C. , Taeyoung Lee

Fluidic locomotion of flapping Micro Aerial Vehicles (MAVs) can be very complex, particularly when the rules from insect flight dynamics (fast flapping dynamics and light wings) are not applicable. In these situations, widely used averaging…

Systems and Control · Electrical Eng. & Systems 2021-10-05 Paul Ghanem , Yunus Bicer , Deniz Erdogmus , Alireza Ramezani

This research endeavors to design the perching maneuver and control in ornithopter robots. By analyzing the dynamic interplay between the robot's flight dynamics, feedback loops, and the environmental constraints, we aim to advance our…

Systems and Control · Electrical Eng. & Systems 2026-01-26 C. Ruiz , J. Á. Acosta

Flapping-wing insects, birds, and robots are thought to offset the high power cost of oscillatory wing motion by using elastic elements for energy storage and return. Insects possess highly resilient elastic regions in their flight anatomy…

Biological Physics · Physics 2020-12-11 James Lynch , Jeffrey Gau , Simon Sponberg , Nick Gravish

Estimating the direction of ambient fluid flow is key for many flying or swimming animals and robots, but can only be accomplished through indirect measurements and active control. Recent work with tethered flying insects indicates that…

Systems and Control · Electrical Eng. & Systems 2022-04-20 Floris van Breugel