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Microswimmers in suspension exhibit collective swimming behaviour, forming various self-organised structures including ordered, aggregated, and turbulent-like structures. When mixed with passive particles phase-separation is known to occur,…

Fluid Dynamics · Physics 2026-03-26 Alexander Chamolly , Takuji Ishikawa

We demonstrate with experiments and simulations how microscopic self-propelled particles navigate through environments presenting complex spatial features, which mimic the conditions inside cells, living organisms and future lab-on-a-chip…

Biological Physics · Physics 2011-09-22 Giovanni Volpe , Ivo Buttinoni , Dominik Vogt , Hans-Juergen Kuemmerer , Clemens Bechinger

A striking feature of the collective behavior of spherical microswimmers is that for sufficiently strong self-propulsion they phase-separate into a dense cluster coexisting with a low-density dis- ordered surrounding. Extending our previous…

Soft Condensed Matter · Physics 2016-10-31 Johannes Blaschke , Maurice Maurer , Karthik Menon , Andreas Zöttl , Holger Stark

We consider the dynamics of active nematics droplets on flat surfaces, based on the continuum hydrodynamic theory. We investigate a wide range of dynamical regimes as a function of the activity and droplet size on surfaces characterized by…

Soft Condensed Matter · Physics 2023-09-11 Rodrigo C. V. Coelho , Hélio R. J. C. Figueiredo , Margarida M. Telo da Gama

We show that hydrodynamic theories of polar active matter generically possess inhomogeneous traveling solutions. We introduce a unifying dynamical-system framework to establish the shape of these intrinsically nonlinear patterns, and show…

We introduce a minimal model for a collection of self-propelled apolar active particles, also called as `active nematic', on a two-dimensional substrate and study the order-disorder transition with the variation of density. The particles…

Statistical Mechanics · Physics 2015-09-18 Rakesh Das , Manoranjan Kumar , Shradha Mishra

A wide range of living and artificial active matter exists in close contact with substrates and under strong confinement, where in addition to dipolar active stresses, quadrupolar active stresses can become important. Here, we numerically…

Soft Condensed Matter · Physics 2024-04-23 Salik A. Sultan , Mehrana R. Nejad , Amin Doostmohammadi

An interacting pair of chemotactic (anti-chemotactic) active colloids, that can rotate their axes of self-propulsion to align {parallel (anti-parallel)} to a chemical gradient, shows dynamical behaviour that varies from bound states to…

Biological Physics · Physics 2025-10-08 Suropriya Saha , Sriram Ramaswamy , Ramin Golestanian

Biological and artificial microswimmers often self-propel in external flows of vortical nature; relevant examples include algae in small-scale ocean eddies, spermatozoa in uterine peristaltic flows and bacteria in microfluidic devices. A…

Biological Physics · Physics 2022-11-14 Ivan Tanasijevic , Eric Lauga

Suspensions of swimming micro-organisms are known to undergo intricate collective dynamics as a result of hydrodynamic and collision interactions. Micro-swimmers, such as bacteria and micro-algae, naturally live and have evolved in complex…

Fluid Dynamics · Physics 2025-10-31 Yasser Almoteri , Enkeleida Lushi

The field of active nematics has traditionally employed descriptions based on dipolar activity, with interactions that align along a single axis. However, it has been theoretically predicted that interactions with a substrate, prevalent in…

We developed a mechanical model of spermatozoal swimming in bulk suspensions. We traced the spatiotemporal elastohydrodynamic interactions and found that spermatozoa engaged in self-organisation: flagellar undulatory motion generated a…

Fluid Dynamics · Physics 2023-02-22 Nanami Taketoshi , Toshihiro Omori , Takuji Ishikawa

Active processes drive and guide biological dynamics across scales -- from subcellular cytoskeletal remodelling, through tissue development in embryogenesis, to population-level bacterial colonies expansion. In each of these, biological…

One objective of active matter science is to unveil principles by which chaotic microscale dynamics could be transformed into useful work. A nematic liquid crystal environment offers a number of possibilities, one of which is a directional…

Soft Condensed Matter · Physics 2021-11-02 Mojtaba Rajabi , Hend Baza , Hao Wang , Oleg D. Lavrentovich

Collective motion is a phenomenon observed across length scales in nature, from bacterial swarming and tissue migration to the flocking of animals. The mechanisms underlying this behavior vary significantly depending on the biological…

Soft Condensed Matter · Physics 2025-01-15 Urvi Mahendra Bora , Mohd Suhail Rizvi

In active nematic liquid crystals activity is able to drive chaotic spatiotemporal flows referred to as active turbulence. Active turbulence has been characterized through theoretical and experimental work as a low Reynolds number…

Soft Condensed Matter · Physics 2022-11-30 C. Rorai , F. Toschi , I. Pagonabarraga

We use continuum simulations to study the impact of anisotropic hydrodynamic friction on the emergent flows of active nematics. We show that, depending on whether the active particles align with or tumble in their collectively self-induced…

Soft Condensed Matter · Physics 2020-03-16 Kristian Thijssen , Luuk Metselaar , Julia M. Yeomans , Amin Doostmohammadi

Growing tissue and bacterial colonies are active matter systems where cell divisions and cellular motion generate active stress. Although they operate in the non-equilibrium regime, these biological systems can form large-scale ordered…

Biological Physics · Physics 2019-05-28 Yusuf Ilker Yaman , Esin Demir , Roman Vetter , Askin Kocabas

The acoustofluidic method holds great promise for manipulating microorganisms. When exposed to the steady vortex structures of acoustic streaming flow, these microorganisms exhibit intriguing dynamic behaviors, such as hydrodynamic trapping…

Fluid Dynamics · Physics 2025-04-25 Xuyang Sun , Wenchang Tan , Yi Man

Motor proteins drive persistent motion and self-organisation of cytoskeletal filaments. However, state-of-the-art microscopy techniques and continuum modelling approaches focus on large length and time scales. Here, we perform…

Soft Condensed Matter · Physics 2024-06-03 Gerrit Vliegenthart , Arvind Ravichandran , Marisol Ripoll , Thorsten Auth , Gerhard Gompper
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