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Collections of interacting, self-propelled particles have been extensively studied as minimal models of many living and synthetic systems from bird flocks to active colloids. However, the influence of active rotations in the absence of…

Soft Condensed Matter · Physics 2016-11-23 Benjamin C. van Zuiden , Jayson Paulose , William T. M. Irvine , Denis Bartolo , Vincenzo Vitelli

Active fluids, such as cytoskeletal filaments, bacterial colonies and epithelial cell layers, exhibit distinctive orientational coherence, often characterized by nematic order and topological defects. By contrast, little is known about…

Soft Condensed Matter · Physics 2021-04-07 Mattia Serra , Linnea Lemma , Luca Giomi , Zvonimir Dogic , L. Mahadevan

Active nematics are materials composed of mobile, elongated particles that can transform energy from the environment into a mechanical motion. Current experimental realizations of the active nematics are of biological origin and include…

Soft Condensed Matter · Physics 2024-06-05 Andrey Sokolov , Jaideep Katuri , Juan J. de Pablo , Alexey Snezhko

Active nematics are dense systems of rodlike particles that consume energy to drive motion at the level of the individual particles. They exist in natural systems like biological tissues and artificial materials such as suspensions of…

Soft Condensed Matter · Physics 2023-12-14 Yunrui Li , Zahra Zarei , Phu N. Tran , Yifei Wang , Aparna Baskaran , Seth Fraden , Michael F. Hagan , Pengyu Hong

We review recent work on active colloids or swimmers, such as self-propelled microorganisms, phoretic colloidal particles, and artificial micro-robotic systems, moving in fluid-like environments. These environments can be water-like and…

Biological Physics · Physics 2016-02-09 Alison E. Patteson , Arvind Gopinath , Paulo E. Arratia

Cell crawling crucially depends on the collective dynamics of the acto-myosin cytoskeleton. However, it remains an open question to what extent cell polarization and persistent motion depend on continuous regulatory mechanisms and…

Biological Physics · Physics 2023-08-11 Fenna Stegemerten , Karin John , Uwe Thiele

The spatiotemporal dynamics in systems of active self-propelled particles is controlled by the propulsion mechanism in combination with various direct interactions, such as steric repulsion, hydrodynamics, and chemical fields. Yet, these…

Soft Condensed Matter · Physics 2024-06-03 Mario Theers , Elmar Westphal , Kai Qi , Roland G. Winkler , Gerhard Gompper

Active (i.e., self-propelled or swimming) particles moving through an isotropic fluid exhibit conventional diffusive behavior. We report anomalous diffusion of an active particle moving in an anisotropic, nematic background. Whilst the…

Soft Condensed Matter · Physics 2016-06-22 John Toner , Hartmut Löwen , Henricus H. Wensink

Active fluids, such as suspensions of microswimmers, are known to self-organize into complex spatio-temporal flow patterns. An intriguing example is mesoscale turbulence, a state of dynamic vortex structures exhibiting a characteristic…

Soft Condensed Matter · Physics 2025-07-02 Henning Reinken , Sebastian Heidenreich , Markus Bär , Sabine H. L. Klapp

We investigate theoretically the collective dynamics of a suspension of low Reynolds number swimmers that are confined to two dimensions by a thin fluid film. Our model swimmer is characterized by internal degrees of freedom which locally…

Biological Physics · Physics 2015-05-19 M. Leoni , T. B. Liverpool

Microswimming cells and robots exhibit diverse behaviours due to both their swimming and their environment. One of the core environmental features impacting inertialess swimming is background flows. While the influence of select flows,…

Fluid Dynamics · Physics 2025-10-30 Eamonn A. Gaffney , Kenta Ishimoto , Benjamin J. Walker

We investigate similarities in the micro-structural dynamics between externally driven and actively driven nematics. Walls, lines of strong deformations in the director field, and topological defects are characteristic features of an active…

Soft Condensed Matter · Physics 2015-04-28 Sumesh P. Thampi , Ramin Golestanian , Julia M. Yeomans

Continuum hydrodynamic models of active liquid crystals have been used to describe dynamic self-organising systems such as bacterial swarms and cytoskeletal gels. A key prediction of such models is the existence of self-stabilising kink…

Soft Condensed Matter · Physics 2009-11-13 S. A. Edwards , J. M. Yeomans

Micropolar active matter requires for its kinematic description both positional and orientational degrees of freedom. Activity generates dynamic coupling between these kinematic variables that are absent in micropolar passive matter, such…

Soft Condensed Matter · Physics 2022-02-15 Austen Bolitho , Ronojoy Adhikari

Dense, active systems show active turbulence, a state characterised by flow fields that are chaotic, with continually changing velocity jets and swirls. Here we review our current understanding of active turbulence. The development is…

Soft Condensed Matter · Physics 2016-08-03 Sumesh P. Thampi , Julia M. Yeomans

Active nematics are an important new paradigm in soft condensed matter systems. They consist of rod-like components with an internal driving force pushing them out of equilibrium. The resulting fluid motion exhibits chaotic advection, in…

We study a continuum model of an extensile active nematic to show that mesoscale turbulence develops in two stages: (i) ordered regions undergo an intrinsic hydrodynamic instability generating walls, lines of stong bend deformations, (ii)…

Soft Condensed Matter · Physics 2015-06-18 Sumesh P. Thampi , Ramin Golestanian , Julia M. Yeomans

Dynamical clustering represents a characteristic feature of active matter consisting of self-propelled agents that convert energy from the environment into mechanical motion. At the micron scale, typical of overdamped dynamics, particles…

Soft Condensed Matter · Physics 2024-06-04 Lorenzo Caprini , Davide Breoni , Anton Ldov , Christian Scholz , Hartmut Löwen

The study of liquid crystals at equilibrium has led to fundamental insights into the nature of ordered materials, as well as to practical applications such as display technologies. Active nematics are a fundamentally different class of…

Soft Condensed Matter · Physics 2015-08-19 Stephen J. DeCamp , Gabriel S. Redner , Aparna Baskaran , Michael F. Hagan , Zvonimir Dogic

Microorganisms ofter move in confined, disordered environments, where hydrodynamic couplings can modify their transport behavior. Using extensive finite-element simulations, we investigate the dynamics of microswimmers -- modeled as…

Soft Condensed Matter · Physics 2026-03-24 Mirko Residori , Sebastian Aland , Christina Kurzthaler