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Many biological fluids are composed of suspended polymers immersed in a viscous fluid. A prime example is mucus, where the polymers are also known to form a network. While the presence of this microstructure is linked with an overall…

Fluid Dynamics · Physics 2024-10-10 Adam K. Townsend , Eric E. Keaveny

We study a novel phase of active polar fluids, which is characterized by the continuous creation and destruction of dense clusters due to self-sustained turbulence. This state arises due to the interplay of the self-advection of the aligned…

Soft Condensed Matter · Physics 2022-07-12 Vasco M. Worlitzer , Gil Ariel , Avraham Be'er , Holger Stark , Markus Bär , Sebastian Heidenreich

Active matter often simultaneously exhibits different kinds of orientational order and, in many cases of biological interest, undergoes continuous material renewal. In renewing nematopolar fluids we find stable topological strings,…

Soft Condensed Matter · Physics 2026-01-27 Alberto Dinelli , Ludovic Dumoulin , Karsten Kruse

Self-propelled active particles exhibit delayed responses to environmental changes, modulating their propulsion speed through intrinsic sensing and feedback mechanisms. This adaptive behavior fundamentally determines their dynamics and…

Soft Condensed Matter · Physics 2025-05-21 Viktor Holubec , Alexander Fischer , Giovanni Volpe , Frank Cichos

Swimmers and self-propelled particles are physical models for the collective behaviour and motility of a wide variety of living systems, such as bacteria colonies, bird flocks and fish schools. Such artificial active materials are amenable…

Active nematics are the nonequilibrium analog of passive liquid crystals in which anisotropic units consume free energy to drive emergent behavior. Similar to liquid crystal (LC) molecules in displays, ordering and dynamics in active…

Quantitative Methods · Quantitative Biology 2022-10-10 Japinder Nijjer , Mrityunjay Kothari , Changhao Li , Thomas Henzel , Qiuting Zhang , Jung-Shen B. Tai , Shuang Zhou , Sulin Zhang , Tal Cohen , Jing Yan

The persistent dynamics in systems out of equilibrium, particularly those characterized by annihilation and creation of topological defects, is known to involve complicated spatiotemporal processes and is deemed difficult to control. Here…

Soft Condensed Matter · Physics 2022-11-22 Zhi-Feng Huang , Hartmut Löwen , Axel Voigt

We study two microswimmers consisting of a spherical rigid head and a passive elastic tail. In the first one the tail is clamped to the head, and the system oscillates under the action of an external torque. In the second one, head and tail…

Fluid Dynamics · Physics 2017-02-02 Giancarlo Cicconofri , Antonio DeSimone

"Active nematics" are orientationally ordered but apolar fluids composed of interacting constituents individually powered by an internal source of energy. When activity exceeds a system-size dependent threshold, spatially uniform active…

Soft Condensed Matter · Physics 2017-11-08 Richard Green , John Toner , Vincenzo Vitelli

Dynamics of small particles, both living such as swimming bacteria and inanimate, such as colloidal spheres, has fascinated scientists for centuries. If one could learn how to control and streamline their chaotic motion, that would open…

Soft Condensed Matter · Physics 2023-02-08 Oleg D. Lavrentovich

The complex formations exhibited by schooling fish have long been the object of fascination for biologists and physicists. However, the physical and sensory mechanisms leading to organized collective behavior remain elusive. On the physical…

Fluid Dynamics · Physics 2025-07-09 Anand U. Oza , Eva Kanso , Michael J. Shelley

We study numerically the hydrodynamics of a self-propelled particle system, consisting of spherical squirmers sedimented on a flat surface. We observe the emergence of dynamic structures, due to the interplay of particle-particle and…

Soft Condensed Matter · Physics 2019-12-20 Zaiyi Shen , Juho S. Lintuvuori

We theoretically describe the dynamics of swimmer populations confined in thin liquid films. We first demonstrate that hydrodynamic interactions between confined swimmers only depend on their shape and are independent of their specific…

Soft Condensed Matter · Physics 2015-06-11 Tommaso Brotto , Jean-Baptiste Caussin , Eric Lauga , Denis Bartolo

For natural microswimmers, the interplay of swimming activity and external flow can promote robust motion, e.g. propulsion against ("upstream rheotaxis") or perpendicular to the direction of flow. These effects are generally attributed to…

Soft Condensed Matter · Physics 2018-01-30 Jaideep Katuri , William E. Uspal , Juliane Simmchen , Albert Miguel-López , Samuel Sánchez

Collective behaviour in suspensions of microswimmers is often dominated by the impact of long-ranged hydrodynamic interactions. These phenomena include active turbulence, where suspensions of pusher bacteria at sufficient densities exhibit…

Soft Condensed Matter · Physics 2019-08-27 Dora Bardfalvy , Henrik Nordanger , Cesare Nardini , Alexander Morozov , Joakim Stenhammar

We explore the mechanism underlying the anomalous rheology of puller-type microswimmer suspensions through direct hydrodynamic simulations. Puller-type swimmers generate contractile flow fields along their swimming direction, leading to…

Soft Condensed Matter · Physics 2025-03-04 Haruki Hayano , Akira Furukawa

We study the universal behavior of a class of active colloids whose design is inspired by the collective dynamics of natural systems like schools of fish and flocks of birds. These colloids, with off-center repulsive interaction sites,…

Soft Condensed Matter · Physics 2024-10-10 Amir Nourhani

We construct a hydrodynamic theory of noisy, apolar active smectics, in bulk suspension or on a substrate. Our predictions include: quasi-long-ranged smectic order in dimension d = 2, and long- ranged in d = 3, extending previously…

Soft Condensed Matter · Physics 2013-03-19 Tapan Chandra Adhyapak , Sriram Ramaswamy , John Toner

Active materials are capable of converting free energy into directional motion, giving rise to striking dynamical phenomena. Developing a general understanding of their structure in relation to the underlying non-equilibrium physics would…

Soft Condensed Matter · Physics 2018-10-17 Nitin Kumar , Rui Zhang , Juan J. de Pablo , Margaret L. Gardel

We study an active matter system comprised of magnetic microswimmers confined in a microfluidic channel and show that it exhibits a new type of self-organized behavior. Combining analytical techniques and Brownian dynamics simulations, we…

Statistical Mechanics · Physics 2021-02-24 Fanlong Meng , Daiki Matsunaga , Benoît Mahault , Ramin Golestanian