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The propulsion of mammalian spermatozoa relies on the spontaneous periodic oscillation of their flagella. These oscillations are driven internally by the coordinated action of ATP-powered dynein motors that exert sliding forces between…

Fluid Dynamics · Physics 2022-10-13 Chenji Li , Brato Chakrabarti , Pedro Castilla , Achal Mahajan , David Saintillan

Using multiparticle collision dynamics simulations, we investigate the swimming dynamics, orientational behavior, and hydrodynamic interactions of a model swimmer designed to mimic the isolated flagellar apparatus ($FA$) of Chlamydomonas…

Biological Physics · Physics 2025-07-01 Sai Venkata Ramana Ambadipudi , Albert Bae , Azam Gholami

The modeling of the beating of cilia and flagella in fluids is a particularly active field of study, given the biological relevance of these organelles. Various mathematical models have been proposed to represent the nonlinear dynamics of…

Soft Condensed Matter · Physics 2024-12-10 Irene Anello , François Alouges , Antonio De Simone

Many biological microorganisms and artificial microswimmers react to external cues of environmental gradients by changing their swimming directions. We study here the behavior of eukarytic flagellated microswimmers in linear viscosity…

Soft Condensed Matter · Physics 2026-05-29 Shubham Anand , Jens Elgeti , Gerhard Gompper

The motility of microorganisms is influenced greatly by their hydrodynamic interactions with the fluidic environment they inhabit. We show by direct experimental observation of the bi-flagellated alga Chlamydomonas reinhardtii that fluid…

Biological Physics · Physics 2015-11-03 Boyang Qin , Arvind Gopinath , Jing Yang , Jerry P Gollub , Paulo E Arratia

Living creatures exhibit a remarkable diversity of locomotion mechanisms, evolving structures specialised for interacting with their environment. In the vast majority of cases, locomotor behaviours such as flying, crawling, and running, are…

Biological Physics · Physics 2020-07-28 Kirsty Y. Wan

Micro-organisms usually can swim in their liquid environment by flagellar or ciliary beating. In this numerical work, we analyze the influence of flagellar beating on the orbits of a swimming cell in a shear flow. We also calculate the…

Soft Condensed Matter · Physics 2017-11-21 Levan Jibuti , Walter Zimmermann , Salima Rafaï , Philippe Peyla

The present habilitation thesis in theoretical biological physics addresses two central dynamical processes in cells and organisms: (i) active motility and motility control and (ii) self-organized pattern formation. The unifying theme is…

Cell Behavior · Quantitative Biology 2018-03-21 Benjamin M. Friedrich

Microscale fluid flows generated by ensembles of beating eukaryotic flagella are crucial to fundamental processes such as development, motility and sensing. Despite significant experimental and theoretical progress, the underlying physical…

Soft Condensed Matter · Physics 2014-03-11 Douglas R. Brumley , Kirsty Y. Wan , Marco Polin , Raymond E. Goldstein

Micro-organisms propel themselves in viscous environments by the periodic, nonreciprocal beating of slender appendages known as flagella. Active materials have been widely exploited to mimic this form of locomotion. However, the realization…

Soft Condensed Matter · Physics 2024-08-06 Ariel Surya Boiardi , Giovanni Noselli

Flagellar motors enable bacteria to navigate their environments by switching rotation direction in response to external cues with high sensitivity. Previous work suggested that ultrasensitivity of the flagellar motor originates from…

Biological Physics · Physics 2025-02-06 Henry H. Mattingly , Yuhai Tu

Run-and-tumble (RT) motion is commonly observed in flagellated microswimmers, arising from synchronous and asynchronous flagellar beating. One such example is a biflagellated alga, called \textit{Chlamydomonas reinhardtii}. Its flagellar…

Soft Condensed Matter · Physics 2025-10-17 Somnath Paramanick , Umashankar Pardhi , Harsh Soni , Nitin Kumar

The eukaryotic flagellum beats periodically, driven by the oscillatory dynamics of molecular motors, to propel cells and pump fluids. Small, but perceivable fluctuations in the beat of individual flagella have physiological implications for…

Cell Behavior · Quantitative Biology 2015-06-18 Rui Ma , Gary S. Klindt , Ingmar H. Riedel-Kruse , Frank Jülicher , Benjamin M. Friedrich

The physical basis of flagellar and ciliary beating is a major problem in biology which is still far from completely understood. The fundamental cytoskeleton structure of cilia and flagella is the axoneme, a cylindrical array of microtubule…

Biological Physics · Physics 2017-03-10 David Oriola , Hermes Gadêlha , Jaume Casademunt

Cilia and flagella exhibit regular bending waves that perform mechanical work on the surrounding fluid, to propel cellular swimmers and pump fluids inside organisms. Here, we quantify a force-velocity relationship of the beating flagellum,…

Cell Behavior · Quantitative Biology 2016-12-21 Gary S. Klindt , Christian Ruloff , Christian Wanger , Benjamin M. Friedrich

The amoeboid organism, the plasmodium of Physarum polycephalum, behaves on the basis of spatio-temporal pattern formation by local contraction-oscillators. This biological system can be regarded as a reaction-diffusion system which has…

patt-sol · Physics 2009-10-31 H. Yamada , T. Nakagaki , M. Ito

Multi-flagellated bacteria utilize the hydrodynamic interaction between their filamentary tails, known as flagella, to swim and change their swimming direction in low Reynolds number flow. This interaction, referred to as bundling and…

Robotics · Computer Science 2023-07-04 Zhuonan Hao , Sangmin Lim , M. Khalid Jawed

Escherichia coli swims using flagella activated by rotary motors. The direction of rotation of the motors is indirectly regulated by the binding of a single messenger protein. The conformational spread model has been shown to accurately…

Quantitative Methods · Quantitative Biology 2017-03-08 Alberto Pezzotta , Matteo Adorisio , Antonio Celani

Eukaryotes swim with coordinated flagellar (ciliary) beating and steer by fine-tuning the coordination. The model organism for studying flagellate motility, C. reinhardtii (CR), employs synchronous, breast-stroke-like flagellar beating to…

Biological Physics · Physics 2024-06-12 Da Wei , Greta Quaranta , Marie-Eve Aubin-Tam , Daniel S. W. Tam

The beating flagella of the green alga Chlamydomonas reinhardtii play a prominent role in cellular mechanics, enabling cells to both displace and sense surrounding fluid. Specifically, flagellum-induced fluid transport enables microalgae to…

Fluid Dynamics · Physics 2026-03-10 Md Iftekhar Yousuf Emon , Gregorius R. Pradipta , Xiang Cheng , Xin Yong
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