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We present a combined numerical and analytical study of pattern formation in an active system where particles align, possess a density-dependent motility, and are subject to a logistic reaction. This is a model for suspensions of…

Soft Condensed Matter · Physics 2015-06-16 X. Yang , D. Marenduzzo , M. C. Marchetti

Proliferating cell collectives often develop an active growth layer near their boundary that regulates expansion and morphology, as observed in systems ranging from bacterial biofilms to epithelial tissues and tumor spheroids. While such…

Soft Condensed Matter · Physics 2026-04-27 Fidel Álvarez-Murphy , Ignacio Medina , Néstor Sepúlveda , Gustavo Düring

The appearance of emergent symmetries in complex systems with components that can form composite units provides us with opportunities for design and control of exotic phase behaviour, for example by exploiting the dynamical symmetry…

Soft Condensed Matter · Physics 2026-03-24 Jakob Metson , Ramin Golestanian

Phase separation driven by nonequilibrium fluctuations is a hallmark of both living and synthetic active matter. Unlike equilibrium systems, where ordered states arise from the minimization of free energy, active systems are fueled by a…

In this paper, we bring together our efforts in identifying and understanding nonequilibrium phase segregation driven by active processes in the living cell, with special focus on the segregation of cell membrane components driven by active…

Soft Condensed Matter · Physics 2022-05-04 Ajay Bansal , Amit Das , Madan Rao

Due to their remarkable properties, systems that exhibit self-organization of their components resulting from intrinsic microscopic activity have been extensively studied in the last two decades. In a generic class of active matter, the…

Statistical Mechanics · Physics 2019-07-24 Ramin Golestanian

While bacterial swimming has been well characterized in uniform liquid environments, only little is known about how bacteria propagate through complex environments, such as gel-like matrices or porous media that are typically encountered in…

Soft Condensed Matter · Physics 2024-08-06 Agniva Datta , Sönke Beier , Veronika Pfeifer , Robert Großmann , Carsten Beta

Nonreciprocal interactions have recently attracted growing interest in nonequilibrium physics. In particular, breaking action-reaction symmetry has been proposed as a mechanism for collective motion, yet controlled experimental realizations…

We present evidence that the concurrent existence of two populations of particles with different effective diameters is not a prerequisite for the occurrence of anomalous phase behaviors in systems of particles interacting through…

Soft Condensed Matter · Physics 2015-05-20 Santi Prestipino , Franz Saija , Gianpietro Malescio

Motility-induced phase separation (MIPS) is a paradigmatic non-equilibrium transition in active matter, determined by the P\'eclet number and packing fraction. We investigate the single-phase and phase-separated regimes of MIPS using a…

Biological Physics · Physics 2025-10-03 Italo Salas , Francisca Guzmán-Lastra , Denisse Pastén , Ariel Norambuena

Microcolonies are aggregates of a few dozen to a few thousand cells exhibited by many bacteria. The formation of microcolonies is a crucial step towards the formation of more mature bacterial communities known as biofilms, but also marks a…

We study a model of bacterial dynamics where two interacting random walkers perform run-and-tumble motion on a one-dimensional lattice under mutual exclusion and find an exact expression for the probability distribution in the steady state.…

Statistical Mechanics · Physics 2016-06-01 A. B. Slowman , M. R. Evans , R. A. Blythe

Systems driven far from equilibrium may exhibit anomalous density fluctuations: active matter with orientational order display giant density fluctuations at large scale, while systems of interacting particles close to an absorbing phase…

Statistical Mechanics · Physics 2026-03-23 Sara Dal Cengio , Romain Mari , Eric Bertin

Mixtures of active and passive particles are predicted to exhibit a variety of nonequilibrium phases. Here we report a dynamic clustering phase in mixtures of colloids and motile bacteria. We show that colloidal clustering results from a…

Soft Condensed Matter · Physics 2022-05-25 Shreyas Gokhale , Junang Li , Alexandre Solon , Jeff Gore , Nikta Fakhri

Active matter is rapidly becoming a key paradigm of out-of-equilibrium soft matter exhibiting complex collective phenomena, yet the thermodynamics of such systems remain poorly understood. In this letter we study the nonequilbrium…

Statistical Mechanics · Physics 2019-10-30 Emanuele Crosato , Mikhail Prokopenko , Richard E. Spinney

The effect of a diffusivity edge is studied in a system of scalar active matter confined by a periodic potential and driven by an externally applied force. We find that this system shows qualitatively distinct stationary regimes depending…

Statistical Mechanics · Physics 2024-02-27 Jonas Berx , Aritra Bose , Ramin Golestanian , Benoît Mahault

In equilibrium, the physical properties of matter are set by the interactions between the constituents. In contrast, the energy input of the individual components controls the behavior of synthetic or living active matter. Great progress…

Soft Condensed Matter · Physics 2025-08-29 Q. Martinet , Y. Li , A. Aubret , E. Hannezo , J. Palacci

Filamentous cyanobacteria can show fascinating examples of nonequilibrium self-organization, which however are not well-understood from a physical perspective. We investigate the motility and collective organization of colonies of these…

Soft Condensed Matter · Physics 2023-10-20 Mixon K. Faluweki , Jan Cammann , Marco G. Mazza , Lucas Goehring

While the breakdown of the perturbation expansion for the many-electron problem has several formal consequences, here we unveil its physical effect: Flipping the sign of the effective electronic interaction in specific scattering channels.…

Strongly Correlated Electrons · Physics 2020-11-06 M. Reitner , P. Chalupa , L. Del Re , D. Springer , S. Ciuchi , G. Sangiovanni , A. Toschi

Colonies of bacteria endowed with a pili-based self-propulsion machinery are ideal models for investigating the structure and dynamics of active many-particle systems. We study Neisseria gonorrhoeae colonies with a molecular-dynamics-based…

Soft Condensed Matter · Physics 2022-10-18 Kai Zhou , Marc Hennes , Berenike Maier , Gerhard Gompper , Benedikt Sabass