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The origin and early evolution of eukaryotes are one of the major transitions in the evolution of life on earth. One of its most interesting aspects is the emergence of cellular organelles, their dynamics, their functions, and their…

Populations and Evolution · Quantitative Biology 2022-02-18 Parsifal Fidelio Islas-Morales , Luis Felipe Jimenez-Garcia

A first principles approach to the theoretical description of the development of biological forms, from a fertilized egg to a functioning embryo, remains a central challenge to applied physics and theoretical biology. Rather than refer to…

Biological Physics · Physics 2021-04-30 Dominic Gastaldo , David Silverstein

The present view of biological phenomena is based on a biochemical paradigm that development of living organisms is defined by information stored in a molecular form as some genetic code. However, new discoveries indicate that biological…

Other Quantitative Biology · Quantitative Biology 2023-11-14 Yuri K. Shestopaloff

The multifarious internal workings of organisms are difficult to reconcile with a single feature defining a state of being alive. Indeed, definitions of life rely on emergent properties (growth, capacity to evolve, agency) only symptomatic…

Other Quantitative Biology · Quantitative Biology 2023-08-16 Simon Pierce

Physical emergence - crystals, rocks, sandpiles, turbulent eddies, planets, stars - is fundamentally different from biological emergence - amoeba, cells, mice, humans - even though the latter is based in the former. This paper points out…

Biological Physics · Physics 2018-03-29 George Ellis , Jonathan Kopel

This short text summarizes the work in biology proposed in our book, Perspectives on Organisms, where we analyse the unity proper to organisms by looking at it from different viewpoints. We discuss the theoretical roles of biological time,…

Other Computer Science · Computer Science 2019-03-14 Giuseppe Longo , Maël Montévil

The emergence of life from inanimate matter presents a thermodynamic challenge: the Second Law of Thermodynamics dictates a global trend towards disorder, yet life constitutes localized pockets of profound organization. This paper presents…

Statistical Mechanics · Physics 2026-03-17 Shlomo Segal

Research on the birth and evolution of life are reviewed with reference to the maximum entropy production principle (MEPP). It has been shown that this principle is essential for consistent understanding of the birth and evolution of life.…

Biological Physics · Physics 2025-04-22 Yasuji Sawada , Yasukazu Daigaku , Kenji Toma

Life is a complex biological phenomenon represented by numerous chemical, physical and biological processes performed by a biothermodynamic system/cell/organism. Both living organisms and inanimate objects are subject to aging, a biological…

Other Quantitative Biology · Quantitative Biology 2018-01-25 Marko Popovic

The concept of evolutionary development of structures constituted a \emph{real} revolution in biology: it was possible to understand how the very complex structures of life can arise in an out-of-equilibrium system. The investigation of…

Populations and Evolution · Quantitative Biology 2009-08-27 Franco Bagnoli

Due to recent advances in synthetic biology and artificial life, the origin of life is currently a hot topic of research. We review the literature and argue that the two traditionally competing "replicator-first" and "metabolism-first"…

Artificial Intelligence · Computer Science 2013-11-21 Tom Froese , Nathaniel Virgo , Takashi Ikegami

Biological molecules, like all active matter, use free energy to generate force and motion which drive them out of thermal equilibrium, and undergo inherent dynamic interconversion between metastable free energy states separated by levels…

Biological Physics · Physics 2025-08-28 Mark C Leake

The origin of life on Earth via the spontaneous emergence of a protocell prior to Darwinian evolution remains a fundamental open question in physics and chemistry. Here, we develop a conceptual framework based on information theory and…

Populations and Evolution · Quantitative Biology 2025-12-23 Robert G. Endres

Biological systems are influenced by fluid mechanics at nearly all spatiotemporal scales. This broad relevance of fluid mechanics to biology has been increasingly appreciated by engineers and biologists alike, leading to continued expansion…

Fluid Dynamics · Physics 2019-11-27 John O. Dabiri

Newton set the stage for our view of how science should be done. We remain in what I will call the `Newtonian Paradigm' in all of physics, including Newton, Einstein, and Schrodinger. As I will show shortly, Newton invented and bequeathed…

History and Philosophy of Physics · Physics 2013-03-25 Stuart Kauffman

Biological systems reach hierarchical complexity that has no counterpart outside the realm of biology. Undoubtedly, biological entities obey the fundamental physical laws. Can today's physics provide an explanatory framework for…

Statistical Mechanics · Physics 2022-10-12 Yuri I. Wolf , Mikhail I. Katsnelson , Eugene V. Koonin

Living systems self-organize in ways that conventional physical frameworks-based on forces, energies, and continuous fields-cannot fully capture. Processes like gene regulation and cellular decision-making involve rule-based logic and…

Other Quantitative Biology · Quantitative Biology 2025-11-18 Hyun Youk

It is probably not a coincidence that two of the pioneers of thermodynamics, Helmholtz and Mayer, were physicians. Thermodynamics studies the transformations of energy, and such transformations ceaselessly take place in all living systems…

Biological Physics · Physics 2007-05-23 Shahar Dolev , Avshalom C. Elitzur

The improbability of a spontaneously generated self-assembling molecule has suggested that life began with a set of simpler, collectively replicating elements, such as an enclosed autocatalytic set of polymers (or protocell). Since…

Adaptation and Self-Organizing Systems · Physics 2013-08-26 Liane Gabora

The living organism is considered as an open system, whereas Prigogine's approach to the thermodynamics of such systems is used. The approach allows one to formulate the law of individual growth and development (ontogenesis) of the living…

Biological Physics · Physics 2026-03-12 Alexei A. Zotin , Vladimir N. Pokrovskii