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When swimming at low Reynolds numbers, inertial effects are negligible and reciprocal movements cannot induce net motion. Instead, symmetry breaking is necessary to achieve net propulsion. Directed swimming can be supported by magnetic…

Soft Condensed Matter · Physics 2026-03-31 Theo Lequy , Andreas M. Menzel

Controlling complex light waves to achieve desired behaviours or characteristics on demand presents a significant challenge. This task becomes even more complicated when manipulating speckled light beams owing to their inherently fuzzy…

In the absence of inertia, a reciprocal swimmer achieves no net motion in a viscous Newtonian fluid. Here, we investigate the ability of a reciprocally actuated particle to translate through a complex fluid that possesses a network using…

Fluid Dynamics · Physics 2015-06-18 David A. Gagnon , Nathan C. Keim , Xiaoning Shen , Paulo E. Arratia

With exquisite precision and reproducibility, cells orchestrate the cooperative action of thousands of nanometer-sized molecular motors to carry out mechanical tasks at much larger length scales, such as cell motility, division and…

Soft Condensed Matter · Physics 2013-01-08 Tim Sanchez , Daniel T. N. Chen , Stephen J. DeCamp , Michael Heymann , Zvonimir Dogic

The directed self-assembly of colloidal nanoparticles (NPs) using external fields guides the formation of sophisticated hierarchical materials but becomes less effective with decreasing particle size. As an alternative, electron-beam-driven…

Soft Condensed Matter · Physics 2024-09-13 Jonathan G. Raybin , Ethan J. Dunsworth , Veronica Guo , Naomi S. Ginsberg

Peritrichously-flagellated bacteria, such as Escherichia coli, self-propel in fluids by using specialised motors to rotate multiple helical filaments. The rotation of each motor is transmitted to a short flexible segment called the hook…

Biological Physics · Physics 2018-06-07 Emily E. Riley , Debasish Das , Eric Lauga

We introduce a system of light driven microscopic autonomous moving particles that move on a flat surface. The design is simple, yet effective: Micrometer sized objects with wedge shape are produced by photopolymerization, they are covered…

Shape anisotropy of individual building blocks plays a crucial role in creating exotic structures and controlling phase behavior in equilibrium systems. We present a combined experimental and simulation study in which we used light-driven…

Soft Condensed Matter · Physics 2026-05-05 Yogesh Shelke , Anpuj Nair S , Hanumantha Rao Vutukuri

Artificial microswimmers are a new technology with promising microfluidics and biomedical applications, such as directed cargo transport, microscale assembly, and targeted drug delivery. A fundamental barrier to realising this potential is…

Fluid Dynamics · Physics 2018-06-27 Thomas D. Montenegro-Johnson

In equilibrium, the physical properties of matter are set by the interactions between the constituents. In contrast, the energy input of the individual components controls the behavior of synthetic or living active matter. Great progress…

Soft Condensed Matter · Physics 2025-08-29 Q. Martinet , Y. Li , A. Aubret , E. Hannezo , J. Palacci

The transport of motile entities across modulated energy landscapes plays an important role in a range of phenomena in biology, colloidal science and solid-state physics. Here, an easily implementable strategy that allows for the collective…

Soft Condensed Matter · Physics 2020-03-11 Fernando Martínez-Pedrero , Francisco Ortega , Ramón G. Rubio , Carles Calero

In recent years the functionality of synthetic active microparticles has edged even closer to that of their biological counterparts. However, we still lack the understanding needed to recreate at the microscale key features of autonomous…

Soft Condensed Matter · Physics 2024-10-28 Philipp W. A. Schönhöfer , Sharon C. Glotzer

Centrifugation is a widespread laboratory technique used to separate mixtures into fractions characterized by a specific size, weight or density. We demonstrate that centrifugation can be also used to separate swimming cells having…

Soft Condensed Matter · Physics 2014-11-07 Claudio Maggi , Alessia Lepore , Jacopo Solari , Alessandro Rizzo , Roberto Di Leonardo

Liquid-liquid phase separation is important across biology, physics, and materials science. Although usually studied at equilibrium, active components - such as motor proteins, enzymes, and synthetic microswimmers - are increasingly…

Soft Condensed Matter · Physics 2026-01-06 Kan Chang , Yulin Li , Ming Yuan , Masaki Sano , Zhihong You , H. P. Zhang

Active systems comprising micron-sized self-propelling units, also termed microswimmers, are promising candidates for the bottom-up assembly of small structures and reconfigurable materials. Here we leverage field-driven colloidal assembly…

Soft Condensed Matter · Physics 2024-12-24 Carolina van Baalen , Laura Alvarez , Robert Style , Lucio Isa

Biofilms are spatially organized microorganism colonies embedded in a self-produced matrix, conferring to the microbial community resistance to environmental stresses. Motile bacteria have been observed swimming in the matrix of pathogenic…

Quantitative Methods · Quantitative Biology 2022-05-24 Guillaume Ravel , Michel Bergmann , Alain Trubuil , Julien Deschamps , Romain Briandet , Simon Labarthe

Active colloids are microscopic particles, which self-propel through viscous fluids by converting energy extracted from their environment into directed motion. We first explain how articial microswimmers move forward by generating…

Soft Condensed Matter · Physics 2016-05-12 Andreas Zöttl , Holger Stark

The run-and-tumble dynamics of bacteria, as exhibited by \textit{E. coli}, offers a simple experimental realization of non-Brownian, yet diffusive, particles. Here we present some analytic and numerical results for models of the ideal…

Statistical Mechanics · Physics 2009-08-12 J. Tailleur , M. E. Cates

The natural habitats of microorganisms in the human microbiome and ocean and soil ecosystems are full of colloids and macromolecules, which impart non-Newtonian flow properties drastically affecting the locomotion of swimming…

Soft Condensed Matter · Physics 2022-04-01 Shashank Kamdar , Seunghwan Shin , Lorraine F. Francis , Xinliang Xu , Xiang Cheng

Motile microorganisms, like bacteria and algae, unify abilities like self-propulsion, autonomous navigation, and decision-making on the micron scale. While recent breakthroughs have led to the creation of synthetic microswimmers and…

Soft Condensed Matter · Physics 2026-02-06 Aritra K. Mukhopadhyay , Ran Niu , Linhui Fu , Kai Feng , Christopher Fujta , Qiang Zhao , Jinping Qu , Benno Liebchen
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