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We investigate theoretically the collective dynamics of a suspension of low Reynolds number swimmers that are confined to two dimensions by a thin fluid film. Our model swimmer is characterized by internal degrees of freedom which locally…

Biological Physics · Physics 2015-05-19 M. Leoni , T. B. Liverpool

Self-propelled colloids (swimmers) in confining geometries follow trajectories determined by hydrodynamic interactions with the bounding surfaces. However, typically these interactions are ignored or truncated to lowest order. We…

Many eukaryotic microorganisms propelled by multiple flagella can swim very rapidly with distinct gaits. Here, we model a three-dimensional mutiflagellate swimming strategy, resembling the microalgae, and investigate the effects of…

Soft Condensed Matter · Physics 2023-10-17 Shiyuan Hu , Fanlong Meng

The motility of microorganisms is influenced greatly by their hydrodynamic interactions with the fluidic environment they inhabit. We show by direct experimental observation of the bi-flagellated alga Chlamydomonas reinhardtii that fluid…

Biological Physics · Physics 2015-11-03 Boyang Qin , Arvind Gopinath , Jing Yang , Jerry P Gollub , Paulo E Arratia

Biological and artificial microswimmers often have to propel through a variety of environments, ranging from heterogeneous suspending media to strong geometrical confinement. Under confinement, local flow fields generated by microswimmers,…

Soft Condensed Matter · Physics 2024-06-04 Florian A. Overberg , Gerhard Gompper , Dmitry A. Fedosov

We use a three-bead-spring model to investigate the dynamics of bi-flagellate micro-swimmers near a surface. While the primary dynamics and scattering are governed by geometric-dependent direct contact, the fluid flows generated by the…

Biological Physics · Physics 2017-08-17 Enkeleida Lushi , Vasily Kantsler , Raymond E. Goldstein

Active fluids can develop spontaneous flow instabilities and complex patterns. However, spatio-temporal control of active particles has remained challenging, despite its relevance in biological and applied contexts. Here, we harnessed…

Soft Condensed Matter · Physics 2025-09-23 Isabelle Eisenmann , Marco Vona , Nicolas Desprat , Takuji Ishikawa , Eric Lauga , Raphaël Jeanneret

Several micro-organisms, such as bacteria, algae, or spermatozoa, use flagella or cilia to swim in a fluid, while many other micro-organisms instead use ample shape deformation, described as amoeboid, to propel themselves by either crawling…

Biological Physics · Physics 2016-09-19 Hao Wu , A. Farutin , W. -F. Hu , M. Thiébaud , S. Rafaï , P. Peyla , M. -C. Lai , C. Misbah

Due to its great efficiency and maneuverability, undulatory swimming is the predominant form of locomotion in aquatic vertebrates. A myriad of animals of different species and sizes oscillate their bodies to propel themselves in aquatic…

Fluid Dynamics · Physics 2026-01-26 Jesús Sánchez-Rodríguez , Christophe Raufaste , Mederic Argentina

Spermatozoa flagella are known to synchronize when swimming in close proximity. We use a model consisting of two-dimensional sheets propagating transverse waves of displacement to demonstrate that fluid forces lead to such synchronization…

Fluid Dynamics · Physics 2011-08-30 Gwynn J. Elfring , Eric Lauga

We present in this fluid dynamics video a novel experimental setup with self-propelled swimmers on a free surface. The swimmers, modeled as flexible thin filaments, are subjected to external electromagnetic forcing driving a propagating…

Fluid Dynamics · Physics 2014-11-04 Sophie Ramananarivo , Benjamin Thiria , Ramiro Godoy-Diana

We present the first time-resolved measurements of the oscillatory velocity field induced by swimming unicellular microorganisms. Confinement of the green alga C. reinhardtii in stabilized thin liquid films allows simultaneous tracking of…

Fluid Dynamics · Physics 2015-05-19 Jeffrey S. Guasto , Karl A. Johnson , J. P. Gollub

We study the dynamics of a prolate spheroidal microswimmer in Poiseuille flow for different flow geometries. When moving between two parallel plates or in a cylindrical microchannel, the swimmer performs either periodic swinging or periodic…

Soft Condensed Matter · Physics 2013-01-24 Andreas Zöttl , Holger Stark

Active matter comprises individually driven units that convert locally stored energy into mechanical motion. Interactions between driven units lead to a variety of non-equilibrium collective phenomena in active matter. One of such phenomena…

Soft Condensed Matter · Physics 2021-06-15 Mingji Huang , Wensi Hu , Siyuan Yang , Quan-Xing Liu , H. P. Zhang

In a recent letter (Friedrich et al., Phys. Rev. Lett. 109:138102, 2012), a minimal model swimmer was proposed that propels itself at low Reynolds numbers by a revolving motion of a pair of spheres. The motion of the two spheres can…

Cell Behavior · Quantitative Biology 2015-06-12 Katja Polotzek , Benjamin M. Friedrich

The hydrodynamics of a flagellated microorganism is investigated when swimming close to a planar free-slip surface by means of numerical solu- tions of the Stokes equations obtained via a Boundary Element Method. Depending on the initial…

Fluid Dynamics · Physics 2017-03-31 Daniela Pimponi , Mauro Chinappi , Paolo Gualtieri , Carlo Massimo Casciola

Hydrodynamic interaction strongly influences the collective behavior of the microswimmers. With this work, we study the behavior of two hydrodynamically interacting self-propelled chiral swimmers in the low Reynolds number regime,…

Soft Condensed Matter · Physics 2022-11-28 Ruma Maity , P. S. Burada

The behavior of fluid interfaces far from equilibrium plays central roles in nature and in industry. Active swimmers trapped at interfaces can alter transport at fluid boundaries with far reaching implications. Swimmers can become trapped…

Soft Condensed Matter · Physics 2024-02-08 Jiayi Deng , Mehdi Molaei , Nicholas G. Chisholm , Kathleen J. Stebe

Swimming micro-organisms such as flagellated bacteria and sperm cells have fascinating locomotion capabilities. Inspired by their natural motion, there is an ongoing effort to develop artificial robotic nano-swimmers for potential in-body…

Fluid Dynamics · Physics 2023-05-31 Jithu Paul , Yizhar Or , Oleg Gendelman

Cilia and flagella often exhibit synchronized behavior; this includes phase locking, as seen in {\it Chlamydomonas}, and metachronal wave formation in the respiratory cilia of higher organisms. Since the observations by Gray and Rothschild…

Soft Condensed Matter · Physics 2016-05-19 Kirsty Y. Wan , Raymond E. Goldstein