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Microorganisms ofter move in confined, disordered environments, where hydrodynamic couplings can modify their transport behavior. Using extensive finite-element simulations, we investigate the dynamics of microswimmers -- modeled as…

Soft Condensed Matter · Physics 2026-03-24 Mirko Residori , Sebastian Aland , Christina Kurzthaler

Biological microswimmers such as bacteria and sperm cells often encounter complex biological fluid environments. Here we use the well-known squirmer microswimmer model to show the importance of the local fluid microstructure and…

Soft Condensed Matter · Physics 2023-07-19 Andreas Zöttl

We present and analyze a theoretical model for the dynamics and interactions of "capillary surfers," which are millimetric objects that self-propel while floating at the interface of a vibrating fluid bath. In our companion paper [1], we…

Fluid Dynamics · Physics 2024-11-25 Anand U. Oza , Giuseppe Pucci , Ian Ho , Daniel M. Harris

Biological microswimmers are known to navigate upstream of an external flow (positive rheotaxis) in trajectories ranging from linear, spiral to oscillatory. Such rheotaxis stems from the interplay between the motion and complex shapes of…

Soft Condensed Matter · Physics 2022-05-27 Ranabir Dey , Carola M. Buness , Babak Vajdi Hokmabad , Chenyu Jin , Corinna C. Maass

Bacteria exist in a free-swimming state or in a sessile biofilm state. The transition from free-swimming to sessile mode is characterized by changes in gene expression which alter, among others,the mechanics of flagellar motility. In this…

Fluid Dynamics · Physics 2014-05-07 Alan Cheng Hou Tsang , Eva Kanso

Interaction of motile microrganisms with a nearby solid substrate is a well studied phenomenon. However, the effects of hydrodynamic slippage on the substrate have received a little attention. In the present study, within the framework of…

Soft Condensed Matter · Physics 2020-05-05 Antarip Poddar , Aditya Bandopadhyay , Suman Chakraborty

The ability of microswimmers to deploy optimal propulsion strategies is of paramount importance for their locomotory performance and survival at low Reynolds numbers. Although for perfectly spherical swimmers minimum dissipation requires a…

Fluid Dynamics · Physics 2021-07-13 Abdallah Daddi-Moussa-Ider , Babak Nasouri , Andrej Vilfan , Ramin Golestanian

Swimming at a micrometer scale demands particular strategies. Indeed when inertia is negligible as compared to viscous forces (i.e. Reynolds number $Re$ is lower than unity), hydrodynamics equations are reversible in time. To achieve…

Soft Condensed Matter · Physics 2011-02-17 Michaël Garcia , Stefano Berti , Philippe Peyla , Salima Rafaï

Swimming cells and microorganisms must often move though complex fluids that contain an immersed microstructure such as polymer molecules, or filaments. In many important biological processes, such as mammalian reproduction and bacterial…

Fluid Dynamics · Physics 2018-08-06 Arshad Kamal , Eric E Keaveny

Recently, an active microswimmer was constructed where a micron-sized droplet of bromine water was placed into a surfactant-laden oil phase. Due to a bromination reaction of the surfactant at the interface, the surface tension locally…

Soft Condensed Matter · Physics 2016-01-12 Maximilian Schmitt , Holger Stark

We theoretically describe the dynamics of swimmer populations confined in thin liquid films. We first demonstrate that hydrodynamic interactions between confined swimmers only depend on their shape and are independent of their specific…

Soft Condensed Matter · Physics 2015-06-11 Tommaso Brotto , Jean-Baptiste Caussin , Eric Lauga , Denis Bartolo

Cilia and flagella exhibit regular bending waves that perform mechanical work on the surrounding fluid, to propel cellular swimmers and pump fluids inside organisms. Here, we quantify a force-velocity relationship of the beating flagellum,…

Cell Behavior · Quantitative Biology 2016-12-21 Gary S. Klindt , Christian Ruloff , Christian Wanger , Benjamin M. Friedrich

Microswimmers typically operate in complex environments. In biological systems, often diverse species are simultaneously present and interact with each other. Here, we derive a (time-dependent) particle-scale statistical description, namely…

Soft Condensed Matter · Physics 2019-08-13 Christian Hoell , Hartmut Löwen , Andreas M. Menzel

Cilia and flagella are actively bending slender organelles, performing functions such as motility, feeding and embryonic symmetry breaking. We review the mechanics of viscous-dominated microscale flow, including time-reversal symmetry, drag…

Quantitative Methods · Quantitative Biology 2013-09-06 Thomas D. Montenegro-Johnson , Andrew A. Smith , David J. Smith , Daniel Loghin , John R. Blake

Microorganisms are rarely found in Nature swimming freely in an unbounded fluid. Instead, they typically encounter other organisms, hard walls, or deformable boundaries such as free interfaces or membranes. Hydrodynamic interactions between…

Fluid Dynamics · Physics 2013-10-21 Marcelo A. Dias , Thomas R. Powers

Active suspensions encompass a wide range of complex fluids containing microscale energy-injecting particles, such as cells, bacteria or artificially powered active colloids. Because they are intrinsically non-equilibrium, active…

Soft Condensed Matter · Physics 2024-07-09 Zhouyang Ge , Gwynn J. Elfring

Although the motility of the flagellated bacteria, Escherichia coli, has been widely studied, the effect of viscosity on swimming speed remains controversial. The swimming mode of wild-type E.coli is often idealized as a "run-and- tumble"…

Biological Physics · Physics 2018-05-09 Zijie Qu , Fatma Zeynep Temel , Rene Henderikx , Kenneth S. Breuer

We experimentally study a scallop-like swimmer with reciprocally flapping wings in a nearly frictionless, cohesive granular medium consisting of hydrogel spheres. Significant locomotion is found when the swimmer's flapping frequency matches…

Soft Condensed Matter · Physics 2026-04-29 Hongyi Xiao , Jing Wang , Achim Sack , Ralf Stannarius , Thorsten Pöschel

We use particle simulations to reveal two distinct propulsion mechanisms for a scallop-like swimmer to locomote itself in granular media by reciprocally flapping its wings. Based on the discrete element method, we examine the kinematics and…

Soft Condensed Matter · Physics 2025-10-28 Amir Nazemi , Hongyi Xiao

We use numerical simulations to probe the dynamics of concentrated suspensions of spherical microswimmers interacting hydrodynamically. Previous work in the dilute limit predicted orientational instabilities of aligned suspensions for both…

Soft Condensed Matter · Physics 2013-02-12 Arthur A. Evans , Takuji Ishikawa , Takami Yamaguchi , Eric Lauga