Related papers: Flagellar swimmers oscillate between pusher- and p…
We show that activity and broken fore-aft shape symmetry enable microswimmers to cross streamlines in nonuniform shear, a key yet overlooked factor in active cross-stream migration. Using a model of flagellated microswimmers in microchannel…
Using a combination of theory, experiments, and numerical simulations, we investigate the stability of coherent structures in a suspension of strongly aligned active swimmers. We show that a dilute jet of pullers undergoes a pearling…
Spermatozoa self-propel by propagating bending waves along a predominantly active elastic flagellum. The organized structure of the "9 + 2" axoneme is lost in the most-distal few microns of the flagellum, and therefore this region is…
We present an experimental study of capillary surfers, a new fluid-mediated active system that bridges the gap between dissipation- and inertia-dominated regimes. Surfers are wave-driven particles that self-propel and interact on a fluid…
Microswimmers, especially in theoretical treatments, are generally taken to be completely inertia-free, since inertial effects on their motion are typically small and assuming their absence simplifies the problem considerably. Yet in nature…
Suspensions of microswimmers are a rich source of fascinating new fluid mechanics. Recently we predicted the active pipe flow dispersion of gyrotactic microalgae, whose orientation is biased by gravity and flow shear. Analytical theory…
We develop a minimal hydrodynamic model, without an orientational order parameter, for assemblies of contractile swimmers encapsulated in a droplet of a binary-fluid emulsion. Our model uses two coupled scalar order parameters, $\phi$ and…
We numerically study the dynamics of run-and-tumble particles confined in two chambers connected by thin channels. Two dominant dynamical behaviors emerge: (i) an oscillatory pumping state, in which particles periodically fill the two…
The dynamics and motion of multi-ciliated microswimmers with a spherical body and a small number N (with 5 < N < 60) of cilia with length comparable to the body radius, is investigated by mesoscale hydrodynamics simulations. A metachronal…
Many microorganisms take a chiral path while swimming in an ambient uid. In this paper, we study the combined behavior of two chiral swimmers using the well-known squirmer model taking into account chiral asymmetries. In contrast to the…
We survey the theory synchronization in collections of noisy oscillators. This framework is applied to flagellar synchronization by hydrodynamic interactions. The time-reversibility of hydrodynamics at low Reynolds numbers prompts swimming…
Concentrated suspensions of swimming microorganisms and other forms of active matter are known to display complex, self-organized spatio-temporal patterns on scales large compared to those of the individual motile units. Despite intensive…
Most classical work on the hydrodynamics of low-Reynolds-number swimming addresses deterministic locomotion in quiescent environments. Thermal fluctuations in fluids are known to lead to a Brownian loss of the swimming direction. As most…
Bacterial swimming is well characterized in uniform liquids at rest. The natural habitat of bacterial swimmers, however, is often dominated by moving fluids and interfaces, resulting in shear flows that may strongly alter bacterial…
Many microorganisms swim through gels and non-Newtonian fluids in their natural environments. In this paper, we focus on microorganisms which use flagella for propulsion. We address how swimming velocities are affected in nonlinearly…
The design of artificial microswimmers is often inspired by the strategies of natural microorganisms. Many of these creatures exploit the fact that elasticity breaks the time-reversal symmetry of motion at low Reynolds numbers, but this…
In isotropic fluids like water, micrometer-scale swimmers have evolved swim strokes to translate despite their tiny size. As described by Purcell in his Scallop Theorem, reciprocal motions, like those performed by a scallop, cannot drive…
Many microswimmers are inherently chiral, and this chirality can introduce fascinating behaviors in a collection of microswimmers. The dynamics become even more intriguing when two types of microswimmers with distinct chirality are mixed.…
Self-propelled microorganisms, such as unicellular algae or bacteria, swim along their director relative to the fluid velocity. Under a steady shear flow the director rotates in close orbit, a periodic structure that is preserved under an…
Active swimmers are ubiquitous in nature, found in many diverse biological systems ranging from bacteria to vertebrate fish. Of particular importance are sperm cells which are swimmers that are crucial for the survival of many species…