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Chiral active particles, or self-propelled circle swimmers, from sperm cells to asymmetric Janus colloids, form a rich set of patterns, which are different from those seen in linear swimmers. Such patterns have mainly been explored for…

Soft Condensed Matter · Physics 2019-07-24 Demian Levis , Benno Liebchen

The hydrodynamic interactions between a sedimenting microswimmer and a solid wall have ubiquitous biological and technological applications. A plethora of gravity-induced swimming dynamics near a planar no-slip wall provides a platform for…

Soft Condensed Matter · Physics 2025-05-14 Henry Shum , Devenayagam Palaniappan , Yuan-Nan Young

Many eukaryotic cells use the active waving motion of flexible flagella to self-propel in viscous fluids. However, the criteria governing the selection of particular flagellar waveforms among all possible shapes has proved elusive so far.…

Biological Physics · Physics 2013-08-02 Christophe Eloy , Eric Lauga

Enhanced diffusion of passive tracers immersed in active fluids is a universal feature of active fluids and has been extensively studied in recent years. Similar to microrheology for equilibrium complex fluids, the unusual enhanced particle…

Soft Condensed Matter · Physics 2016-11-03 Ou Yang , Yi Peng , Zhengyang Liu , Chao Tang , Xinliang Xu , Xiang Cheng

Self-propelled phoretic swimmers are generally studied in the laminar flow regime, where their low speed renders inertial effects negligible and trajectories highly predictable. This research tackles the challenge of propulsion in the…

Fluid Dynamics · Physics 2026-01-19 Alessandro Foradori , Paolo Bettotti

The present habilitation thesis in theoretical biological physics addresses two central dynamical processes in cells and organisms: (i) active motility and motility control and (ii) self-organized pattern formation. The unifying theme is…

Cell Behavior · Quantitative Biology 2018-03-21 Benjamin M. Friedrich

We investigate a self-organized swimmer at low Reynolds numbers. The microscopic swimmer is composed of three spheres that are connected by two identical active linker arms. Each linker arm contains molecular motors and elastic elements and…

Biological Physics · Physics 2009-09-30 Stefan Gunther , Karsten Kruse

Swimming in circles occurs in a variety of situations at low Reynolds number. Here we propose a simple model for a swimmer that undergoes circular motion, generalising the model of a linear swimmer proposed by Najafi and Golestanian (Phys.…

Soft Condensed Matter · Physics 2012-07-16 Rodrigo Ledesma-Aguilar , Hartmut Loewen , Julia M. Yeomans

The stochastic dynamics of flagellar beating for micro-swimmers, such as flagellated cells, sperms and microalgae, is widely thought to include a feedback mechanism between flagellar shape and the rate of activation/de-activation of the $N…

Soft Condensed Matter · Physics 2026-05-01 F. Fanelli , A. Puglisi

Active matter exhibits various forms of non-equilibrium states in the absence of external forcing, including macroscopic steady-state currents. Such states are often too complex to be modelled from first principles and our understanding of…

Soft Condensed Matter · Physics 2020-09-17 Viktor Škultéty , Cesare Nardini , Joakim Stenhammar , Davide Marenduzzo , Alexander Morozov

Sperm are propelled by bending waves travelling along the flagellum. During steering in gradients of sensory cues, sperm adjust the flagellar beat waveform. Symmetric and asymmetric beat waveforms produce straight and curved swimming paths,…

Microorganism motility often takes place within complex, viscoelastic fluid environments, e.g., sperm in cervicovaginal mucus and bacteria in biofilms. In such complex fluids, strains and stresses generated by the microorganism are stored…

Cells swimming in confined environments are attracted by surfaces. We measure the steady-state distribution of smooth-swimming bacteria (Escherichia coli) between two glass plates. In agreement with earlier studies, we find a strong…

Soft Condensed Matter · Physics 2008-10-02 Allison P. Berke , Linda Turner , Howard C. Berg , Eric Lauga

We use the boundary element method to study the low-Reynolds number locomotion of a spherical model microorganism in a circular tube. The swimmer propels itself by tangen- tial or normal surface motion in a tube whose radius is on the order…

Fluid Dynamics · Physics 2013-06-11 Lailai Zhu , Eric Lauga , Luca Brandt

Microswimmers exhibit noisy circular motion due to asymmetric propulsion mechanisms, their chiral body shape, or by hydrodynamic couplings in the vicinity of surfaces. Here, we employ the Brownian circle swimmer model and characterize…

Soft Condensed Matter · Physics 2019-11-11 Christina Kurzthaler , Thomas Franosch

We run numerical simulations of strongly confined suspensions of model spherical swimmers called "squirmers". Because of the confinement, the Stokeslet dipoles generated by the particles are quickly screened and the far-field flow is…

Fluid Dynamics · Physics 2016-06-22 Jean-Baptiste Delfau , John Molina , Masaki Sano

Natural swimmers usually perform undulations to propel themselves and perform a range of maneuvers. These include various biological species ranging from micro-sized organisms to large-sized fishes that undulate at typical kinematic…

Fluid Dynamics · Physics 2020-07-01 Muhammad Saif Ullah Khalid , Junshi Wang , Haibo Dong , Moubin Liu

We use boundary element simulations to study the interaction of model microswimmers with a neutrally buoyant spherical particle. The ratio of the size of the particle to that of the swimmer is varied from $R^\mathrm{P} / R^\mathrm{S} \ll…

Soft Condensed Matter · Physics 2017-11-01 Henry Shum , Julia M. Yeomans

Flagellated bacteria on nutrient-rich substrates can differentiate into a swarming state and move in dense swarms across surfaces. A recent experiment measured the flow in the fluid around an Escherichia coli swarm (Wu, Hosu and Berg, 2011…

Biological Physics · Physics 2016-07-28 Justas Dauparas , Eric Lauga

Unlike macroscopic swimmers, microswimmers operate in a low-Reynolds-number regime dominated by viscous forces. This paper investigates the controllability of a magnetic microswimmer composed of a spherical magnetic head and an elastic,…

Optimization and Control · Mathematics 2025-11-05 Lucas Palazzolo , Mickaël Binois , Laëtitia Giraldi