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In bacterial chemotaxis, E. coli cells drift up chemical gradients by a series of runs and tumbles. Runs are periods of directed swimming, and tumbles are abrupt changes in swimming direction. Near the beginning of each run, the rotating…

Soft Condensed Matter · Physics 2009-11-07 Thomas R. Powers

Many active biological particles, such as swimming microorganisms or motor-proteins, do work on their environment by going though a periodic sequence of shapes. Interactions between particles can lead to the phase-synchronization of their…

Soft Condensed Matter · Physics 2022-07-26 Brato Chakrabarti , Michael J. Shelley , Sebastian Fürthauer

Efficient mixing of fluids is essential in many practical applications to achieve homogeneity. For microscopic systems, however, both diffusion and turbulence are ineffective methods to achieve chaotic mixing due to the low Reynolds number,…

Soft Condensed Matter · Physics 2025-09-18 Yue Wang , Jonas Berx

Optical vortex traps created from helical modes of light can drive fluid-borne colloidal particles in circular trajectories. Concentric circulating rings of particles formed by coaxial optical vortices form a microscopic Couette cell, in…

Soft Condensed Matter · Physics 2015-06-25 Kosta Ladavac , David G. Grier

Many biological microorganisms and artificial microswimmers react to external cues of environmental gradients by changing their swimming directions. We study here the behavior of eukarytic flagellated microswimmers in linear viscosity…

Soft Condensed Matter · Physics 2026-05-29 Shubham Anand , Jens Elgeti , Gerhard Gompper

Most bacteria swim through fluids by rotating helical flagella which can take one of twelve distinct polymorphic shapes. The most common helical waveform is the "normal" form, used during forward swimming runs. To shed light on the…

Biological Physics · Physics 2015-03-17 Saverio E. Spagnolie , Eric Lauga

It has been a long-standing problem how schooling fish optimize their motion by exploiting the vortices shed by the others. A recent experimental study showed that a pair of fish reduce energy consumption by matching the phases of their…

Biological Physics · Physics 2023-12-05 Susumu Ito , Nariya Uchida

Motivated by recent simulations and by experiments on aggregation of gliding bacteria, we study a model of the collective dynamics of self-propelled hard rods on a substrate in two dimensions. The rods have finite size, interact via…

Soft Condensed Matter · Physics 2009-11-13 Aparna Baskaran , M. Cristina Marchetti

The flow-driven transport of interacting micron-sized particles occurs in many soft matter systems spanning from the translocation of proteins to moving emulsions in microfluidic devices. Here we combine experiments and theory to…

Soft Condensed Matter · Physics 2022-11-16 Dominik Lips , Eric Cereceda-López , Antonio Ortiz-Ambriz , Pietro Tierno , Artem Ryabov , Philipp Maass

Many swimming microorganisms, such as bacteria and sperm, use flexible flagella to move through viscoelastic media in their natural environments. In this paper we address the effects a viscoelastic fluid has on the motion and beating…

Soft Condensed Matter · Physics 2009-11-13 Henry C. Fu , Charles W. Wolgemuth , Thomas R. Powers

We calculate the hydrodynamic flow field generated far from a cilium which is attached to a surface and beats periodically. In the case of two beating cilia, hydrodynamic interactions can lead to synchronization of the cilia, which are…

Biological Physics · Physics 2007-05-23 Andrej Vilfan , Frank Julicher

Microorganisms are rarely found in Nature swimming freely in an unbounded fluid. Instead, they typically encounter other organisms, hard walls, or deformable boundaries such as free interfaces or membranes. Hydrodynamic interactions between…

Fluid Dynamics · Physics 2013-10-21 Marcelo A. Dias , Thomas R. Powers

Eukaryotes swim with coordinated flagellar (ciliary) beating and steer by fine-tuning the coordination. The model organism for studying flagellate motility, C. reinhardtii (CR), employs synchronous, breast-stroke-like flagellar beating to…

Biological Physics · Physics 2024-06-12 Da Wei , Greta Quaranta , Marie-Eve Aubin-Tam , Daniel S. W. Tam

We define a model microswimmer with a variable cycle time, thus allowing the possibility of phase locking driven by hydrodynamic interactions between swimmers. We find that, for extensile or contractile swimmers, phase locking does occur,…

Soft Condensed Matter · Physics 2015-05-13 Victor B. Putz , Julia M. Yeomans

When a millimetric body is placed atop a vibrating liquid bath, the relative motion between the object and interface generates outward propagating waves with an associated momentum flux. Prior work has shown that isolated chiral objects,…

Soft Condensed Matter · Physics 2025-02-28 Jack-William Barotta , Giuseppe Pucci , Eli Silver , Alireza Hooshanginejad , Daniel M. Harris

Recent experiments proposed to use confined bacteria in order to generate flows near surfaces. We develop a mathematical and a computational model of this fluid transport using a linear superposition of fundamental flow singularities. The…

Biological Physics · Physics 2018-02-27 Justas Dauparas , Debasish Das , Eric Lauga

We present experiments demonstrating tube formation in giant unilamellar vesicles that are suspended in a bath of swimming E. coli bacteria. We chose the lipids such that the bacteria have no adhereing interaction with the membrane. The…

Soft Condensed Matter · Physics 2025-09-19 Makarand Diwe , P B Sunil Kumar , Pramod Pullarkat

To initiate tumbling of an E. coli, one of the helical flagella reverses its sense of rotation. It then transforms from its normal form first to the transient semicoiled state and subsequently to the curly-I state. The dynamics of…

Biological Physics · Physics 2016-02-23 Tapan Chandra Adhyapak , Holger Stark

Potentially habitable planets can orbit close enough to their host star that the differential gravity across their diameters can fix the rotation rate at a specific frequency, a process called tidal locking. Tidally locked planets on…

Earth and Planetary Astrophysics · Physics 2017-10-18 Rory Barnes

Hydrodynamics and confinement dominate bacterial mobility near solid or air-water boundaries, causing flagellated bacteria to move in circular trajectories. This phenomenon results from the counter-rotation between the bacterial body and…

Biological Physics · Physics 2018-10-09 George Araujo , Weijie Chen , Sridhar Mani , Jay X. Tang