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Semiflexible polymers such as filamentous actin play a vital role in the mechanical behavior of cells, yet the basic properties of cross-linked F-actin networks remain poorly understood. To address this issue, we have performed numerical…

Soft Condensed Matter · Physics 2009-11-10 D. A. Head , A. J. Levine , F. C. MacKintosh

The mechanical properties of cells are dominated by the cytoskeleton, an interconnected network of long elastic filaments. The connections between the filaments are provided by crosslinking proteins, which constitute, next to the filaments,…

Soft Condensed Matter · Physics 2012-10-09 Claus Heussinger

Living matter moves, deforms, and organizes itself. In cells this is made possible by networks of polymer filaments and crosslinking molecules that connect filaments to each other and that act as motors to do mechanical work on the network.…

Soft Condensed Matter · Physics 2020-12-15 Sebastian Fürthauer , Daniel J Needleman , Michael J. Shelley

Experiments have shown that elasticity of disordered filamentous networks with compliant crosslinks is very different from networks with rigid crosslinks. Here, we model and analyze filamentous networks as a collection of randomly oriented…

Biological Physics · Physics 2015-06-17 A. Sharma , M. Sheinman , K. M. Heidemann , F. C. MacKintosh

The cytoskeleton is an inhomogeneous network of semi-flexible filaments, which are involved in a wide variety of active biological processes. Although the cytoskeletal filaments can be very stiff and embedded in a dense and cross-linked…

Biological Physics · Physics 2019-05-24 Ines Weber , Cécile Appert-Rolland , Grégory Schehr , Ludger Santen

We present a theoretical framework for the linear and nonlinear visco-elastic properties of reversibly crosslinked networks of semiflexible polymers. In contrast to affine models where network strain couples to the polymer end-to-end…

Soft Condensed Matter · Physics 2016-07-06 Jan Plagge , Andreas Fischer , Claus Heussinger

Disordered networks of semiflexible filaments are common support structures in biology. Familiar examples include fibrous matrices in blood clots, bacterial biofilms, and essential components of cells and tissues of plants, animals, and…

Soft Condensed Matter · Physics 2023-09-28 Saamiya Syed , Fred C. MacKintosh , Jordan L. Shivers

Filamentous bio-materials such as fibrin or collagen networks exhibit an enormous stiffening of their elastic moduli upon large deformations. This pronounced nonlinear behavior stems from a significant separation between the stiffnesses…

Soft Condensed Matter · Physics 2019-05-21 Robbie Rens , Carlos Villarroel , Gustavo Düring , Edan Lerner

Intermediate filaments are cytoskeletal proteins that are key regulators of cell mechanics, a role which is intrinsically tied to their hierarchical structure and their unique ability to accommodate large axial strains. However, how the…

Biological Physics · Physics 2019-03-26 Anders Aufderhorst-Roberts , Gijsje H. Koenderink

In cells, cytoskeletal filament networks are responsible for cell movement, growth, and division. Filaments in the cytoskeleton are driven and organized by crosslinking molecular motors. In reconstituted cytoskeletal systems, motor activity…

The cytoskeleton of eukaryotic cells provides mechanical support and governs intracellular transport. These functions rely on the complex mechanical properties of networks of semiflexible protein filaments. Recent theoretical interest has…

Soft Condensed Matter · Physics 2012-11-01 M. Atakhorrami , G. H. Koenderink , J. F. Palierne , F. C. MacKintosh , C. F. Schmidt

Disordered filamentous networks with compliant crosslinks exhibit a low linear elastic shear modulus at small strains, but stiffen dramatically at high strains. Experiments have shown that the elastic modulus can increase by up to three…

Biological Physics · Physics 2015-01-06 Knut M. Heidemann , Abhinav Sharma , Florian Rehfeldt , Christoph F. Schmidt , Max Wardetzky

Previous research on semiflexible polymers including cytoskeletal networks in cells has suggested the existence of distinct regimes of elastic response, in which the strain field is either uniform (affine) or non-uniform (non-affine) under…

Soft Condensed Matter · Physics 2009-11-11 D. A. Head , A. J. Levine , F. C. MacKintosh

Inspired by the ubiquity of composite filamentous networks in nature we investigate models of biopolymer networks that consist of interconnected floppy and stiff filaments. Numerical simulations carried out in three dimensions allow us to…

Biomolecules · Quantitative Biology 2015-05-19 Elisabeth M. Huisman , Claus Heussinger , Cornelis Storm , Gerard T. Barkema

We introduce a mathematical model for the mechanical behaviour of the eukaryotic cell cytoskeleton. This discrete model involves a regular array of pre-stressed protein filaments that exhibit resistance to enthalpic stretching, joined at…

Quantitative Methods · Quantitative Biology 2023-09-06 J. Köry , N. A. Hill , X. Y. Luo , P. S. Stewart

Cytoskeletal networks are the main actuators of cellular mechanics, and a foundational example for active matter physics. In cytoskeletal networks, motion is generated on small scales by filaments that push and pull on each other via…

Soft Condensed Matter · Physics 2022-10-05 Sebastian Fürthauer , Michael J. Shelley

The actin cytoskeleton in living cells has many types of crosslinkers. The mechanical interplay between these different crosslinker types is an open issue in cytoskeletal mechanics. We develop a framework to study the cooperativity and…

Soft Condensed Matter · Physics 2015-05-28 Moumita Das , D. A. Quint , J. M. Schwarz

Collagen is the most abundant extracellular-network-forming protein in animal biology and is important in both natural and artificial tissues, where it serves as a material of great mechanical versatility. This versatility arises from its…

Soft Condensed Matter · Physics 2009-07-13 D. Vader , A. Kabla , D. Weitz , L. Mahadevan

Motivated by recent experiments showing nonlinear elasticity of in vitro networks of the biopolymer actin cross-linked with filamin, we present an effective medium theory of flexibly cross-linked stiff polymer networks. We model such…

Soft Condensed Matter · Physics 2008-10-19 C. P. Broedersz , C. Storm , F. C. MacKintosh

Networks of elastic beams can deform either by stretching or bending of their members. The primary mode of deformation (bending or stretching) crucially depends on the specific details of the network architecture. In order to shed light on…

Soft Condensed Matter · Physics 2016-09-23 Gérald Gurtner , Marc Durand
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