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Both natural and artificial small-scale swimmers may often self-propel in environments subject to complex geometrical constraints. While most past theoretical work on low-Reynolds number locomotion addressed idealised geometrical…

Fluid Dynamics · Physics 2017-11-16 Alexander Chamolly , Takuji Ishikawa , Eric Lauga

Microorganisms living in microfluidic environments often form multi-species swarms, where they can leverage collective motions to achieve enhanced transport and spreading. Nevertheless, there is a general lack of physical understandings of…

Fluid Dynamics · Physics 2022-02-14 Bryce Palmer , Wen Yan , Tong Gao

The effect of gravity on the collective motion of living microswimmers, such as bacteria and micro-algae, is pivotal to unravel not only bio-convection patterns but also the settling of bacterial biofilms on solid surfaces. In this work, we…

Soft Condensed Matter · Physics 2025-04-07 C. Miguel Barriuso G. , Horacio Serna , Ignacio Pagonabarraga , Chantal Valeriani

We study, numerically, the collective dynamics of self-rotating nonaligning particles by considering a monolayer of spheres driven by constant clockwise or counterclockwise torques. We show that hydrodynamic interactions alter the emergence…

Soft Condensed Matter · Physics 2016-03-24 Kyongmin Yeo , Enkeleida Lushi , Petia M. Vlahovska

In the presented work we study, by means of numerical simulations, the behaviour of a suspension of active ring polymers in the bulk and under lateral confinement. When changing the separation between the confining planes and the polymers'…

Soft Condensed Matter · Physics 2023-07-31 Juan Pablo Miranda-López , Emanuele Locatelli , Chantal Valeriani

Organisms often swim through density stratified fluids. In this Letter, we investigate the dynamics of small active particles swimming in density gradients and report theoretical evidence of taxis as a result of density stratification…

Soft Condensed Matter · Physics 2023-08-02 Vaseem A. Shaik , Gwynn J. Elfring

Self-propelled particles can spontaneously form dense phases from a dilute suspension in a process referred to as motility-induced phase separation. The properties of the out-of-equilibrium structures that are formed are governed by the…

Soft Condensed Matter · Physics 2016-09-08 Sven Erik Ilse , Christian Holm , Joost de Graaf

We present a hydrodynamic study of a monolayer of squirmer model microswimmers confined to a boundary by strong gravity using the simulation method of multi-particle collision dynamics. The squirmers interact with each other via their…

Soft Condensed Matter · Physics 2019-07-23 Jan-Timm Kuhr , Felix Rühle , Holger Stark

The "squirmer model" is a classical hydrodynamic model for the motion of interfacially-driven microswimmers, such as self-phoretic colloids or volvocine green algae. To date, most studies using the squirmer model have considered spherical…

Soft Condensed Matter · Physics 2023-10-31 Ruben Poehnl , William E. Uspal

Motility-Induced Phase Separation (MIPS) is a distinctive phenomenon in active matter that arises from its inherent non-equilibrium nature. Despite recent progress in understanding MIPS in dry active systems, it has been debated whether…

Soft Condensed Matter · Physics 2025-10-01 Tingtao Zhou , John F. Brady

Collections of interacting, self-propelled particles have been extensively studied as minimal models of many living and synthetic systems from bird flocks to active colloids. However, the influence of active rotations in the absence of…

Soft Condensed Matter · Physics 2016-11-23 Benjamin C. van Zuiden , Jayson Paulose , William T. M. Irvine , Denis Bartolo , Vincenzo Vitelli

We investigate a self-organized swimmer at low Reynolds numbers. The microscopic swimmer is composed of three spheres that are connected by two identical active linker arms. Each linker arm contains molecular motors and elastic elements and…

Biological Physics · Physics 2009-09-30 Stefan Gunther , Karsten Kruse

Understanding the hydrodynamics of microswimmers in viscoelastic fluids and confined environments is crucial for interpreting their behaviour in natural settings and designing synthetic microswimmers for practical applications like cargo…

Fluid Dynamics · Physics 2025-01-28 KVS Chaithanya , Sumesh P. Thampi

Understanding how microscopic motility shapes emergent collective behaviors is a challenging task in active matter, especially when self-propulsion is regulated by external cues or via quorum-sensing interactions. To address this problem,…

Statistical Mechanics · Physics 2026-04-13 Alberto Dinelli , Pietro Luigi Muzzeddu

Swimming cells and microorganisms must often move though complex fluids that contain an immersed microstructure such as polymer molecules, or filaments. In many important biological processes, such as mammalian reproduction and bacterial…

Fluid Dynamics · Physics 2018-08-06 Arshad Kamal , Eric E Keaveny

We show that a two-dimensional system of flocking microswimmers interacting hydrodynamically can be expressed using a Hamiltonian formalism. The Hamiltonian depends strictly on the angles between the particles and their swimming…

Soft Condensed Matter · Physics 2023-05-23 Yuval Shoham , Naomi Oppenheimer

We simulate clustering, phase separation and hexatic ordering in a monolayered suspension of active squirming disks subject to an attractive Lennard-Jones-like pairwise interaction potential, taking hydrodynamic interactions between the…

Soft Condensed Matter · Physics 2016-03-31 Ricard Matas Navarro , Suzanne Fielding

Self-propelled particles can navigate complex environments, including viscous fluid interfaces with curved geometries. In this work, we study the emergent dynamics of a suspension of self-propelled particles confined to a stationary curved…

Fluid Dynamics · Physics 2026-04-17 Yuzhu Chen , Vishal P. Patil , David Saintillan

Hydrodynamic interactions driven by particle activity are ubiquitous in active colloidal systems. Although these interactions are strongly influenced by the interfacial actuation mechanism and geometry of the swimming particles, theoretical…

Soft Condensed Matter · Physics 2025-06-17 Anson G. Thambi , William E. Uspal

Similarly to their biological counterparts, suspensions of chemically-active autophoretic swimmers exhibit nontrivial dynamics involving self-organization processes as a result of inter-particle interactions. Using a kinetic model for a…

Soft Condensed Matter · Physics 2022-12-01 Tullio Traverso , Sebastien Michelin