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Hydrodynamic interactions between swimming or flying organisms can lead to complex flows on the scale of the group. These emergent fluid dynamics are often more complex than a linear superposition of individual organism flows, especially at…

Fluid Dynamics · Physics 2024-12-23 Nina Mohebbi , Joonha Hwang , Matthew K. Fu , John O. Dabiri

We derive from first principles a three-dimensional theory of self-propelled particle swarming in a viscous fluid environment. Our model predicts emergent collective behavior that depends critically on fluid opacity, mechanism of…

Soft Condensed Matter · Physics 2026-05-12 Yao-Li Chuang , M. R. D'Orsogna , T. Chou

A three-dimensional (3D) scanning velocimetry system is developed to quantify the 3D configurations of particles and their surrounding volumetric, three-component velocity fields. The approach uses a translating laser sheet to rapidly scan…

Fluid Dynamics · Physics 2021-08-04 Matt K. Fu , Isabel A. Houghton , John O. Dabiri

Active particles, which interact hydrodynamically, display a remarkable variety of emergent collective phenomena. We use squirmers to model spherical microswimmers and explore the collective behavior of thousands of them under the influence…

Fluid Dynamics · Physics 2017-10-31 Jan-Timm Kuhr , Johannes Blaschke , Felix Rühle , Holger Stark

Differential buoyancy surface sources in the ocean may induce a density-driven flow that joins faster flow components to create a multi-scale, 3D flow. Potential temperature and salinity are active tracers that determine the ocean's…

Geophysics · Physics 2022-08-10 Ori Saporta-Katz , Edriss S. Titi , Hezi Gildor , Vered Rom-Kedar

Bacteria in biofilms form complex structures and can collectively migrate within mobile aggregates, which is referred to as swarming. This behavior is influenced by a combination of various factors, including morphological characteristics…

Soft Condensed Matter · Physics 2024-01-19 Bohan Wu-Zhang , Dmitry A. Fedosov , Gerhard Gompper

Bacterial swimming is well characterized in uniform liquids at rest. The natural habitat of bacterial swimmers, however, is often dominated by moving fluids and interfaces, resulting in shear flows that may strongly alter bacterial…

Suspensions of unicellular microswimmers such as flagellated bacteria or motile algae exhibit spontaneous density heterogeneities at large enough concentrations. Based on the relative location of the biological actuation appendages i.e.…

Soft Condensed Matter · Physics 2017-11-27 Fabian Jan Schwarzendahl , Marco G. Mazza

Self-propelled colloids constitute an important class of intrinsically non-equilibrium matter. Typically, such a particle moves ballistically at short times, but eventually changes its orientation, and displays random-walk behavior in the…

Soft Condensed Matter · Physics 2019-05-27 Jochen Arlt , Vincent A Martinez , Angela Dawson , Teuta Pilizota , Wilson C K Poon

Organisms often swim through density stratified fluids. In this Letter, we investigate the dynamics of small active particles swimming in density gradients and report theoretical evidence of taxis as a result of density stratification…

Soft Condensed Matter · Physics 2023-08-02 Vaseem A. Shaik , Gwynn J. Elfring

We investigate theoretically the collective dynamics of a suspension of low Reynolds number swimmers that are confined to two dimensions by a thin fluid film. Our model swimmer is characterized by internal degrees of freedom which locally…

Biological Physics · Physics 2015-05-19 M. Leoni , T. B. Liverpool

It has been proposed that biologically generated turbulence plays an important role in material transport and ocean mixing. Both experimental and numerical studies have reported evidence of the non-negligible mixing by moderate Reynolds…

Fluid Dynamics · Physics 2023-10-06 Xinyu Si , Lei Fang

Microswimming cells and robots exhibit diverse behaviours due to both their swimming and their environment. One of the core environmental features impacting inertialess swimming is background flows. While the influence of select flows,…

Fluid Dynamics · Physics 2025-10-30 Eamonn A. Gaffney , Kenta Ishimoto , Benjamin J. Walker

The presence of stratified layer in atmosphere and ocean leads to buoyant vertical motions, commonly referred to as plumes. It is important to study the mixing dynamics of a plume at a local scale in order to model their evolution and…

Fluid Dynamics · Physics 2016-11-04 Harish N Mirajkar , Sridhar Balasubramanian

Systems of motile microorganisms exhibit a multitude of collective phenomena, including motility-induced phase separation and turbulence. Sensing of the environment and adaptation of movement plays an essential role in the emergent…

Soft Condensed Matter · Physics 2024-06-26 Segun Goh , Elmar Westphal , Roland G. Winkler , Gerhard Gompper

We investigate the fluid dynamics of brine shrimp larvae swimming in this gallery of fluid motion video. Time resolved particle image velocimetry was performed using nano-particles as seeding material to measure the time dependent velocity…

Fluid Dynamics · Physics 2011-10-19 Brennan Johnson , Deborah Garrity , Lakshmi Prasad Dasi

We report on an investigation of the vertical transport of tracer particles released within a shallow, continuously-forced flow by means of numerical simulations. The investigation is motivated by the shallow flows encountered in many…

Motile bacteria play essential roles in biology that rely on their dynamic behaviors, including their ability to navigate, interact, and self-organize. However, bacteria dynamics on fluid interfaces are not well understood. Swimmers…

Biological Physics · Physics 2023-05-02 Jiayi Deng , Mehdi Molaei , Nicholas G. Chisholm , Kathleen J. Stebe

Competition between biological species in marine environments is affected by the motion of the surrounding fluid. An effective 2D compressibility can arise, for example, from the convergence and divergence of water masses at the depth at…

Populations and Evolution · Quantitative Biology 2019-01-24 Abigail Plummer , Roberto Benzi , David R. Nelson , Federico Toschi

We conduct experiments with flexible swimmers to address the impact of fluid viscoelasticity on their locomotion. The swimmers are composed of a magnetic head actuated in rotation by a frequency-controlled magnetic field and a flexible tail…

Fluid Dynamics · Physics 2013-03-19 Julian Espinosa-Garcia , Eric Lauga , Roberto Zenit
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