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We explore hydrodynamic interactions between microswimmers and corrugated, or rough, surfaces, as found often in biological systems and microfluidic devices. Using the Lorentz reciprocal theorem for viscous flows we derive exact expressions…

Soft Condensed Matter · Physics 2022-08-10 Christina Kurzthaler , Howard A. Stone

The asymmetries that arise when a mixing layer involves two miscible fluids of differing densities are investigated using incompressible (low-speed) direct numerical simulations. The simulations are performed in the temporal configuration…

Fluid Dynamics · Physics 2021-01-08 Jon R. Baltzer , Daniel Livescu

Microswimmers typically operate in complex environments. In biological systems, often diverse species are simultaneously present and interact with each other. Here, we derive a (time-dependent) particle-scale statistical description, namely…

Soft Condensed Matter · Physics 2019-08-13 Christian Hoell , Hartmut Löwen , Andreas M. Menzel

Collective behavior models, such as aggregation and flocking, usually assume self-propelled robots that can directly execute their desired speed and direction of motion without fundamental constraints. However, autonomous sailing robots…

Multiagent Systems · Computer Science 2026-05-28 Pranav Kedia , Aaron Gan , Hannah J. Williams , Andreagiovanni Reina , Heiko Hamann

We consider the stirring of an inviscid fluid caused by the locomotion of bodies through it. The swimmers are approximated by non-interacting cylinders or spheres moving steadily along straight lines. We find the displacement of fluid…

Fluid Dynamics · Physics 2010-07-22 Jean-Luc Thiffeault , Stephen Childress

Biological microswimmers such as bacteria and sperm cells often encounter complex biological fluid environments. Here we use the well-known squirmer microswimmer model to show the importance of the local fluid microstructure and…

Soft Condensed Matter · Physics 2023-07-19 Andreas Zöttl

One of the principal mechanisms by which surfaces and interfaces affect microbial life is by perturbing the hydrodynamic flows generated by swimming. By summing a recursive series of image systems we derive a numerically tractable…

Soft Condensed Matter · Physics 2016-10-12 Arnold J. T. M. Mathijssen , Amin Doostmohammadi , Julia M. Yeomans , Tyler N. Shendruk

Recent research has shown that motile cells can adapt their mode of propulsion to the mechanical properties of the environment in which they find themselves--crawling in some environments while swimming in others. The latter can involve…

Biological Physics · Physics 2018-05-24 Hao Wu , Marco Avila Ponce de Leon , Hans G. Othmer

Confined suspensions of active particles show peculiar dynamics characterized by wall accumulation, as well as upstream swimming, centerline depletion and shear-trapping when a pressure-driven flow is imposed. We use theory and numerical…

Fluid Dynamics · Physics 2023-07-19 Barath Ezhilan , David Saintillan

We consider two systems of active swimmers moving close to a solid surface, one being a living population of wild-type \textit{E. coli} and the other being an assembly of self-propelled Au-Pt rods. In both situations, we have identified two…

We present a hydrodynamic study of a monolayer of squirmer model microswimmers confined to a boundary by strong gravity using the simulation method of multi-particle collision dynamics. The squirmers interact with each other via their…

Soft Condensed Matter · Physics 2019-07-23 Jan-Timm Kuhr , Felix Rühle , Holger Stark

We examine the vertical mixing induced by the swimming of microorganisms at low Reynolds and P\'eclet numbers in a stably stratified ocean, and show that the global contribution of oceanic microswimmers to vertical mixing is negligible. We…

Fluid Dynamics · Physics 2014-06-16 Gregory L. Wagner , William R. Young , Eric Lauga

We explore the mechanism underlying the anomalous rheology of puller-type microswimmer suspensions through direct hydrodynamic simulations. Puller-type swimmers generate contractile flow fields along their swimming direction, leading to…

Soft Condensed Matter · Physics 2025-03-04 Haruki Hayano , Akira Furukawa

Controlling bacterial surface adhesion and subsequent biofilm formation in fluid systems is crucial for the safety and efficacy of medical and industrial processes. Here, we theoretically examine the transport of bacteria close to surfaces,…

Soft Condensed Matter · Physics 2025-07-08 Edwina F. Yeo , Benjamin J. Walker , Philip Pearce , Mohit P. Dalwadi

Swimming of a sphere in a viscous incompressible fluid is studied on the basis of the Navier-Stokes equations for wave-type distortions of the spherical shape. At sizable values of the dimensionless scale number the mean swimming velocity…

Fluid Dynamics · Physics 2019-02-20 B. U. Felderhof , R. B. Jones

Micro-swimmers can serve as cargo carriers that move deep inside complex flow networks. When a school collectively entrains the surrounding fluid, their transport capacity can be enhanced. This effect is quantified with good agreement…

Soft Condensed Matter · Physics 2021-08-26 Chenyu Jin , Yibo Chen , Corinna C. Maass , Arnold J. T. M. Mathijssen

The actomyosin cytoskeleton, a protein assembly comprising actin fibers and the myosin molecular motor, drives various cellular dynamics through contractile force generation at high densities. However, the relationship between the density…

Soft Condensed Matter · Physics 2025-01-07 Tomoka Kashiwabara , Tatsuya Fukuyama , Yusuke T. Maeda

In the present paper the behavior of a single artificial microswimmer is addressed, namely an active droplet moving by Marangoni flow. The non-uniform surface tension distribution underlying the propulsion mechanism of the droplet, is…

Computational Physics · Physics 2017-09-13 Laura Stricker

Microswimmers (planktonic microorganisms or artificial active particles) immersed in a fluid interact with the ambient flow, altering their trajectories. In surface gravity waves, a common goal for microswimmers is vertical migration (e.g.,…

Soft Condensed Matter · Physics 2022-03-01 Kunlin Ma , Nimish Pujara , Jean-Luc Thiffeault

Active matter comprises individually driven units that convert locally stored energy into mechanical motion. Interactions between driven units lead to a variety of non-equilibrium collective phenomena in active matter. One of such phenomena…

Soft Condensed Matter · Physics 2021-06-15 Mingji Huang , Wensi Hu , Siyuan Yang , Quan-Xing Liu , H. P. Zhang