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To date, the mechanisms underlying the diversity of the emergent patterns of collective motion in locust hopper bands remain to be unveiled. This study investigates the role of speed heterogeneity in the emergence of the most common…

Neurons and Cognition · Quantitative Biology 2023-03-01 Adrian Bach

In this paper, we explore experimentally the phase behavior of a dense active suspension of self- propelled colloids. In addition to a solid-like and a gas-like phase observed for high and low densities, a novel cluster phase is reported at…

Soft Condensed Matter · Physics 2015-06-04 I. Theurkauff , C. Cottin-Bizonne , J. Palacci , C. Ybert , L. Bocquet

Animal movements have been related to optimal foraging strategies where self-similar trajectories are central. Most of the experimental studies done so far have focused mainly on fitting statistical models to data in order to test for…

Populations and Evolution · Quantitative Biology 2015-06-23 Octavio Miramontes , Og DeSouza , Leticia Ribeiro Paiva , Alessandra Marins , Sirio Orozco

The spontaneous emergence of collective flows is a generic property of active fluids and often leads to chaotic flow patterns characterised by swirls, jets, and topological disclinations in their orientation field. However, the ability to…

Soft Condensed Matter · Physics 2017-03-07 Tyler N. Shendruk , Amin Doostmohammadi , Kristian Thijssen , Julia M. Yeomans

We introduce a stochastic agent-based model for the flocking dynamics of self-propelled particles that exhibit velocity-alignment interactions with neighbours within their field of view. The stochasticity in the dynamics of the model arises…

Statistical Mechanics · Physics 2019-07-24 Trilochan Bagarti , Shakti N. Menon

We derive from first principles a three-dimensional theory of self-propelled particle swarming in a viscous fluid environment. Our model predicts emergent collective behavior that depends critically on fluid opacity, mechanism of…

Soft Condensed Matter · Physics 2026-05-12 Yao-Li Chuang , M. R. D'Orsogna , T. Chou

Aerial displays of starlings (Sturnus vulgaris) at their communal roosts are complex: thousands of individuals form multiple flocks which are continually changing shape and density, while splitting and merging. To understand these complex…

Quantitative Methods · Quantitative Biology 2011-08-26 H. Hildenbrandt , C. Carere , C-K. Hemelrijk

Many systems involve numerous interacting parts and the whole system can have properties that the individual parts do not. I take this novelty as the defining characteristic of an emergent property. Other characteristics associated with…

History and Philosophy of Physics · Physics 2025-11-05 Ross H. McKenzie

Birds in a flock move in a correlated way, resulting in large polarization of velocities. A good understanding of this collective behavior exists for linear motion of the flock. Yet observing actual birds, the center of mass of the group…

Systems of motile microorganisms exhibit a multitude of collective phenomena, including motility-induced phase separation and turbulence. Sensing of the environment and adaptation of movement plays an essential role in the emergent…

Soft Condensed Matter · Physics 2024-06-26 Segun Goh , Elmar Westphal , Roland G. Winkler , Gerhard Gompper

We consider a self-propelled particle system which has been used to describe certain types of collective motion of animals, such as fish schools and bird flocks. Interactions between particles are specified by means of a pairwise potential,…

Analysis of PDEs · Mathematics 2013-03-05 J. A. Carrillo , S. Martin , V. Panferov

Active particles such as swimming bacteria or self-propelled colloids are known to spontaneously organize into fascinating large-scale dynamic structures. The emergence of these collective states from the motility pattern of the individual…

Soft Condensed Matter · Physics 2019-11-20 Hamid Karan , Gerardo E. Pradillo , Petia M. Vlahovska

We propose a novel computational method to extract information about interactions among individuals with different behavioral states in a biological collective from ordinary video recordings. Assuming that individuals are acting as finite…

Multiagent Systems · Computer Science 2017-10-03 Hiroki Sayama , Farnaz Zamani Esfahlani , Ali Jazayeri , J. Scott Turner

Polar active matter - including animal herds, aggregates of motile cells and active colloids - often forms coordinated migration patterns, such as flocking. This orderly motion can be disrupted by full-integer topological defects…

Soft Condensed Matter · Physics 2026-05-26 Luiza Angheluta , Anna Lång , Emma Lång , Stig Ove Bøe

Mixtures of active and passive particles are predicted to exhibit a variety of nonequilibrium phases. Here we report a dynamic clustering phase in mixtures of colloids and motile bacteria. We show that colloidal clustering results from a…

Soft Condensed Matter · Physics 2022-05-25 Shreyas Gokhale , Junang Li , Alexandre Solon , Jeff Gore , Nikta Fakhri

Collective behavior is all around us, from flocks of birds to schools of fish. These systems are immensely complex, which makes it pertinent to study their behavior through minimal models. We introduce such a minimal model for cohesive and…

Soft Condensed Matter · Physics 2025-01-29 Jeanine Shea , Holger Stark

Rotating clusters or vortices are formations of agents that rotate around a common center. These patterns may be found in very different contexts: from swirling fish to surveillance drones. Here, we propose a minimal model for…

Adaptation and Self-Organizing Systems · Physics 2023-05-16 Julia Cantisán , Jesús M. Seoane , Miguel A. F. Sanjuán

Self-propelled particles can spontaneously form dense phases from a dilute suspension in a process referred to as motility-induced phase separation. The properties of the out-of-equilibrium structures that are formed are governed by the…

Soft Condensed Matter · Physics 2016-09-08 Sven Erik Ilse , Christian Holm , Joost de Graaf

Recent investigations have provided important insights into the complex structure and dynamics of collectively moving flocks of living organisms. Two intriguing observations are, scale-free correlations in the velocity fluctuations, in the…

Biological Physics · Physics 2022-03-22 Kunal Bhattacharya , Abhijit Chakraborty

The complex interactions underlying collective motion in biological systems give rise to emergent behaviours such as flocking, sorting, and cooperative transport. These dynamics often involve species with different motilities coordinating…

Soft Condensed Matter · Physics 2025-06-19 Laura Alvarez , Elena Sesé-Sansa , Demian Levis , Ignacio Pagonabarraga , Lucio Isa