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Active systems comprised of self-propelled units show fascinating transitions from Brownian-like dynamics to collective coherent motion. Swirling of swimming bacteria is a spectacular example. This study demonstrates that a nematic liquid…

Several active organisms in nature tend to reside as a community in a viscous fluid medium. We analyze the variation of swimming characteristics of an active swimmer present in a dilute and disperse suspension, modeled as an effective…

Soft Condensed Matter · Physics 2021-10-04 Abyaya Dhar , P. S. Burada , G. P. Raja Sekhar

Swimming micro-organisms such as flagellated bacteria and sperm cells have fascinating locomotion capabilities. Inspired by their natural motion, there is an ongoing effort to develop artificial robotic nano-swimmers for potential in-body…

Fluid Dynamics · Physics 2023-05-31 Jithu Paul , Yizhar Or , Oleg Gendelman

In this review we discuss the recent progress in the simulation of soft active matter systems and in particular the hydrodynamics of microswimmers using the method of multiparticle collision dynamics, which solves the hydrodynamic flows…

Soft Condensed Matter · Physics 2020-09-03 Andreas Zöttl

Turbulence is ubiquitous, from oceanic currents to small-scale biological and quantum systems. Self-sustained turbulent motion in microbial suspensions presents an intriguing example of collective dynamical behavior amongst the simplest…

We show that a two-dimensional system of flocking microswimmers interacting hydrodynamically can be expressed using a Hamiltonian formalism. The Hamiltonian depends strictly on the angles between the particles and their swimming…

Soft Condensed Matter · Physics 2023-05-23 Yuval Shoham , Naomi Oppenheimer

Biological microswimmers often encounter deformable boundaries in physiological conditions; for instance, the viscoelastic walls of reproductive tract during migration of spermatozoa, or host tissue during early bacterial biofilm formation.…

Soft Condensed Matter · Physics 2025-08-07 Smita S. Sontakke , Aneesha Kajampady , Mohd Suhail Rizvi , Ranabir Dey

We derive from first principles a three-dimensional theory of self-propelled particle swarming in a viscous fluid environment. Our model predicts emergent collective behavior that depends critically on fluid opacity, mechanism of…

Soft Condensed Matter · Physics 2026-05-12 Yao-Li Chuang , M. R. D'Orsogna , T. Chou

Microorganisms and synthetic microswimmers often encounter complex environments consisting of networks of obstacles embedded into viscous fluids. Such settings include biological media, such as mucus with filamentous networks, as well as…

The microalga Chlamydomonas Reinhardtii (CR) is used here as a model system to study the effect of complex environments on the swimming of micro-organisms. Its motion can be modelled by a run and tumble mechanism so that it describes a…

Soft Condensed Matter · Physics 2019-03-27 Marvin Brun-Cosme-Bruny , Eric Bertin , Benoît Coasne , Philippe Peyla , Salima Rafaï

After colliding with a surface, microswimmers reside there during the detention time. They accumulate and may form complex structures such as biofilms. We introduce a general framework to calculate the distribution of detention times using…

Soft Condensed Matter · Physics 2015-07-21 Konstantin Schaar , Andreas Zöttl , Holger Stark

We identify the presence of a continuum percolation transition in model suspensions of pusher-type microswimmers. The clusters dynamically aggregate and disaggregate resulting from a competition of attractive and repulsive hydrodynamic and…

Soft Condensed Matter · Physics 2019-08-29 Fabian Jan Schwarzendahl , Marco G. Mazza

Geometric confinements are frequently encountered in the biological world and strongly affect the stability, topology, and transport properties of active suspensions in viscous flow. Based on a far-field analytical model, the…

Cooperative motion in biological microswimmers is crucial for their survival as it facilitates adhesion to surfaces, formation of hierarchical colonies, efficient motion, and enhanced access to nutrients. Synthetic microswimmers currently…

Soft Condensed Matter · Physics 2022-04-12 Stefania Ketzetzi , Melissa Rinaldin , Pim Dröge , Joost de Graaf , Daniela J. Kraft

The hydrodynamic interactions of a suspension of self-propelled particles are studied using a direct numerical simulation method which simultaneously solves for the host fluid and the swimming particles. A modified version of the "Smoothed…

Soft Condensed Matter · Physics 2013-01-15 John J. Molina , Yasuya Nakayama , Ryoichi Yamamoto

In this paper we systematically derive a fourth-order continuum theory capable of reproducing mesoscale turbulence in a three-dimensional suspension of microswimmers. We start from overdamped Langevin equations for a generic microscopic…

Soft Condensed Matter · Physics 2018-06-28 Henning Reinken , Sabine H. L. Klapp , Markus Bär , Sebastian Heidenreich

Microswimmers are active particles of microscopic size that self-propel by setting the surrounding fluid into motion. According to the kind of far-field fluid flow that they induce, they are classified into pushers and pullers. Many studies…

Soft Condensed Matter · Physics 2019-08-01 Giorgio Pessot , Hartmut Löwen , Andreas M. Menzel

The general problem of tracer diffusion in non-equilibrium baths is important in a wide range of systems, from the cellular level to geographical lengthscales. In this paper, we revisit the archetypical example of such a system: a…

Soft Condensed Matter · Physics 2023-11-08 Henrik Nordanger , Alexander Morozov , Joakim Stenhammar

Using a fluid-particle dynamics approach, we numerically study the effects of hydrodynamic interactions on the collective dynamics of active suspensions within a simple model for bacterial motility: each microorganism is modeled as a…

Soft Condensed Matter · Physics 2015-06-22 Akira Furukawa , Davide Marenduzzo , Michael E Cates

Hydrodynamic interactions are crucial for determining the cooperative behavior of microswimmers at low Reynolds numbers. Here we provide a comprehensive analysis of the scaling and strength of the interactions in the case of a pair of…

Soft Condensed Matter · Physics 2021-02-11 Sebastian Ziegler , Thomas Scheel , Maxime Hubert , Jens Harting , Ana-Sunčana Smith