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Suspensions of self-propelled particles, such as swimming micro-organisms, are known to undergo complex dynamics as a result of hydrodynamic interactions. This fluid dynamics video presents a numerical simulation of such a suspension, based…

Fluid Dynamics · Physics 2009-10-20 David Saintillan , Amir Alizadeh Pahlavan

Microswimmers typically operate in complex environments. In biological systems, often diverse species are simultaneously present and interact with each other. Here, we derive a (time-dependent) particle-scale statistical description, namely…

Soft Condensed Matter · Physics 2019-08-13 Christian Hoell , Hartmut Löwen , Andreas M. Menzel

Many biological fluids are composed of suspended polymers immersed in a viscous fluid. A prime example is mucus, where the polymers are also known to form a network. While the presence of this microstructure is linked with an overall…

Fluid Dynamics · Physics 2024-10-10 Adam K. Townsend , Eric E. Keaveny

Microswimmers exhibit an intriguing, highly-dynamic collective motion with large-scale swirling and streaming patterns, denoted as active turbulence -- reminiscent of classical high-Reynolds-number hydrodynamic turbulence. Various…

Soft Condensed Matter · Physics 2024-06-03 Kai Qi , Elmar Westphal , Gerhard Gompper , Roland G. Winkler

Microorganisms can preferentially orient and move along gradients of a chemo-attractant (i.e., chemotax) while colonies of many microorganisms can collectively undergo complex dynamics in response to chemo-attractants that they themselves…

Biological Physics · Physics 2013-10-30 Enkeleida Lushi , Raymond E. Goldstein , Michael J. Shelley

Microswimmers, especially in theoretical treatments, are generally taken to be completely inertia-free, since inertial effects on their motion are typically small and assuming their absence simplifies the problem considerably. Yet in nature…

Soft Condensed Matter · Physics 2016-11-08 Jayant Pande , Kristina Pickl , Oleg Trosman , Ulrich Rüde , Ana-Sunčana Smith

We theoretically describe the dynamics of swimmer populations confined in thin liquid films. We first demonstrate that hydrodynamic interactions between confined swimmers only depend on their shape and are independent of their specific…

Soft Condensed Matter · Physics 2015-06-11 Tommaso Brotto , Jean-Baptiste Caussin , Eric Lauga , Denis Bartolo

Collective behaviour in suspensions of microswimmers is often dominated by the impact of long-ranged hydrodynamic interactions. These phenomena include active turbulence, where suspensions of pusher bacteria at sufficient densities exhibit…

Soft Condensed Matter · Physics 2019-08-27 Dora Bardfalvy , Henrik Nordanger , Cesare Nardini , Alexander Morozov , Joakim Stenhammar

When attracted by a stimulus (e. g. light), microswimmers can build up very densely at a constriction and thus cause clogging. The micro-alga \textit{Chlamydomonas Reinhardtii} is used here as a model system to study this phenomenon. Its…

Soft Condensed Matter · Physics 2020-01-07 Marvin Brun-Cosme-Bruny , Vincent Borne , Sylvain Faure , Bertrand Maury , Philippe Peyla , Salima Rafai

Suspensions of swimming particles exhibit complex collective behaviors driven by hydrodynamic interactions, showing persistent large-scale flows and long-range correlations. While heavily studied, it remains unclear how such structures…

Soft Condensed Matter · Physics 2025-05-27 Bryce Palmer , Scott Weady , Michael O'Brien , Blakesley Burkhart , Michael J. Shelley

Natural habitats of most living microorganisms are distinguished by a complex structure often formed by a porous medium such as soil. The dynamics and transport properties of motile microorganisms are strongly affected by crowded and…

Soft Condensed Matter · Physics 2023-06-23 Florian von Rüling , Liubov Bakhchova , Ulrike Steinmann , Alexey Eremin

Many microorganisms swim in a highly heterogeneous environment with obstacles such as fibers or polymers. To better understand how this environment affects microorganism swimming, we study propulsion of a cylinder or filament in a fluid…

Fluid Dynamics · Physics 2016-04-13 Nguyenho Ho , Karin Leiderman , Sarah D. Olson

Self-propelled particles with hydrodynamic interactions (microswimmers) have previously been shown to produce long-range ordering phenomena. Many theoretical explanations for these collective phenomena are connected to instabilities in the…

Soft Condensed Matter · Physics 2017-05-03 Yuzhou Qian , Peter R. Kramer , Patrick T. Underhill

We investigate the dynamics of model microswimmers under confinement, in cylindrical geometries, by means of three dimensional direct numerical calculations with fully resolved hydrodynamics. Such swimmers are known to show collective…

Soft Condensed Matter · Physics 2017-09-26 Norihiro Oyama , John Jairo Molina , Ryoichi Yamamoto

We study different types of microswimmers moving in channels with varying cross section and thereby interacting hydrodynamically with the channel walls. Starting from the Smoluchowski equation for a dilute suspension, for which interactions…

Soft Condensed Matter · Physics 2017-05-24 Paolo Malgaretti , Holger Stark

Many microorganisms live and evolve in complex fluids. Examples include mammalian spermatozoa in cervical mucus, worms (e.g., \textit{C. elegans}) in wet soil, and bacteria (e.g., \textit{H. pylori}) in our stomach lining. Due to the…

Fluid Dynamics · Physics 2022-10-21 Paulo E. Arratia

Transport of material across liquid interfaces is ubiquitous for living cells and is also a crucial step in drug delivery and in many industrial processes. The fluids that are present on either side of the interfaces will usually have…

Soft Condensed Matter · Physics 2023-05-24 Chao Feng , John J. Molina , Matthew S. Turner , Ryoichi Yamamoto

Microswimming cells and robots exhibit diverse behaviours due to both their swimming and their environment. One of the core environmental features impacting inertialess swimming is background flows. While the influence of select flows,…

Fluid Dynamics · Physics 2025-10-30 Eamonn A. Gaffney , Kenta Ishimoto , Benjamin J. Walker

Biological microswimmers such as bacteria and sperm cells often encounter complex biological fluid environments. Here we use the well-known squirmer microswimmer model to show the importance of the local fluid microstructure and…

Soft Condensed Matter · Physics 2023-07-19 Andreas Zöttl

Hydrodynamical interactions of active micro-particles are pervasive in our planet's fluid environments. Hence, understanding the interactions of these self-propelled particles is essential for science and engineering. In this paper the…

Soft Condensed Matter · Physics 2022-12-28 Yonatan Ashenafi