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Flapping insects are remarkably agile fliers, adapted to a highly turbulent environment. We present a series of high resolution numerical simulations of a bumblebee interacting with turbulent inflow. We consider both tethered and free…

Insects use flight muscles attached at the base of the wings to produce impressive wing flapping frequencies. The maximum power output of these flight muscles is insufficient to maintain such wing oscillations unless there is good elastic…

Fluid Dynamics · Physics 2016-07-20 Yangyang Huang , Eva Kanso

In swarms of flying insects, the motions of individuals are largely uncoordinated with those of their neighbors, unlike the highly ordered motion of bird flocks. However, it has been observed that insects may transiently form pairs with…

We derive a new method to infer from data the out-of-equilibrium alignment dynamics of collectively moving animal groups, by considering the maximum entropy distribution consistent with temporal and spatial correlations of flight direction.…

We present the hydrodynamic theory of coherent collective motion ("flocking") at a solid-liquid interface, and many of its predictions for experiment. We find that such systems are stable, and have long-range orientational order, over a…

Soft Condensed Matter · Physics 2022-01-05 Niladri Sarkar , Abhik Basu , John Toner

Systems of motile microorganisms exhibit a multitude of collective phenomena, including motility-induced phase separation and turbulence. Sensing of the environment and adaptation of movement plays an essential role in the emergent…

Soft Condensed Matter · Physics 2024-06-26 Segun Goh , Elmar Westphal , Roland G. Winkler , Gerhard Gompper

In some physical and biological swarms, agents effectively move and interact along curved surfaces. The associated constraints and symmetries can affect collective-motion patterns, but little is known about pattern stability in the presence…

Adaptation and Self-Organizing Systems · Physics 2020-09-02 Jason Hindes , Victoria Edwards , Sayomi Kamimoto , George Stantchev , Ira B. Schwartz

Demonstrating and quantifying the respective roles of social interactions and external stimuli governing fish dynamics is key to understanding fish spatial distribution. If seminal studies have contributed to our understanding of fish…

Biological Physics · Physics 2015-06-03 Manuela Capello , Marc Soria , Pascal Cotel , Jean-Louis Deneubourg , Laurent Dagorn

Birds rely on active high-acceleration morphing and flapping to navigate complex airflows, but they can also maintain stable fixed-wing postures under persistent atmospheric disturbances. Here, we show that avian wings exhibit aerodynamic…

Fluid Dynamics · Physics 2025-07-22 Lunbing Chen , Suyang Qin , Jinpeng Huang , Yufei Yin , Yang Xiang , Hong Liu

Animals having a trend to align their velocities to an average of their neighbors' may flock as illustrated by the Vicsek model and its variants. If, in addition, they feel a systematic contrarian trend, the result may be a time periodic…

Statistical Mechanics · Physics 2019-02-06 L L Bonilla , C. Trenado

We classify and predict the asymptotic dynamics of a class of swarming models. The model consists of a conservation equation in one dimension describing the movement of a population density field. The velocity is found by convolving the…

Populations and Evolution · Quantitative Biology 2015-05-13 A. J. Leverentz , C. M. Topaz , A. J. Bernoff

In this paper we consider interacting particle systems which are frequently used to model collective behavior in animal swarms and other applications. We study the stability of orientationally aligned formations called flock solutions, one…

Dynamical Systems · Mathematics 2014-03-31 J. A. Carrillo , Y. Huang , S. Martin

In this paper, we proposed a stochastic model which describes two species of particles moving in counterflow. The model generalizes the theoretical framework describing the transport in random systems since particles can work as mobile…

Soft Condensed Matter · Physics 2017-08-09 Eduardo Velasco Stock , Roberto da Silva , Henrique Almeida Fernandes

We investigate the stability of self-propelled particle flocks in the Taylor-Green vortex, a steady vortical flow. We consider a model where particles align themselves to a combination of the orientation and the acceleration of particles…

Biological Physics · Physics 2016-07-13 Andrew W. Baggaley

Soaring migrants exploit columns of rising air (thermals) to cover large distances with minimal energy. Employing social information while locating thermals may be beneficial, but examining collective movements in wild migrants has been a…

Biological Physics · Physics 2018-07-17 Andrea Flack , Mate Nagy , Wolfgang Fiedler , Iain D. Couzin , Martin Wikelski

We consider the collective behaviour of active particles that locally align with their neighbours. Agent-based simulation models have previously shown that in one dimension, these particles can form into a flock that maintains its stability…

Statistical Mechanics · Physics 2018-12-26 Eoin Ó Laighléis , Martin R. Evans , Richard A. Blythe

The cohesive collective motion (flocking, swarming) of autonomous agents is ubiquitously observed and exploited in both natural and man-made settings, thus, minimal models for its description are essential. In a model with continuous space…

Statistical Mechanics · Physics 2015-01-12 Illes J. Farkas , Jeromos Kun , Yi Jin , Gaoqi He , Mingliang Xu

Stability of flapping flight, a natural requirement for flying insects, is one of the major challenges for designing micro aerial vehicles (MAVs). To better understand how a flying insect could stabilize itself during hover, we have…

Fluid Dynamics · Physics 2018-12-06 Chao Zhang , Tyson L. Hedrick , Rajat Mittal , Yijin Mao

As a rough model for the collective motions of cells and organisms we develop here the statistical mechanics of swarms of self-propelled particles. Our approach is closely related to the recently developed theory of active Brownian motion…

Statistical Mechanics · Physics 2007-05-23 Werner Ebeling , Udo Erdmann

Collective motion in animal groups, such as swarms of insects, flocks of birds, and schools of fish, are some of the most visually striking examples of emergent behavior. Empirical analysis of these behaviors in experiment or computational…

Soft Condensed Matter · Physics 2021-03-16 Kevin Schultz , Marisel Villafane-Delgado , Elizabeth P. Reilly , Grace M. Hwang , Anshu Saksena
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