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From bird flocks to fish schools, animal groups often seem to react to environmental perturbations as if of one mind. Most studies in collective animal behaviour have aimed to understand how a globally ordered state may emerge from simple…

Populations and Evolution · Quantitative Biology 2014-10-10 Andrea Cavagna , Alessio Cimarelli , Irene Giardina , Giorgio Parisi , Raffaele Santagati , Fabio Stefanini , Massimiliano Viale

As the constituents of a flock are polar, one expects a fore-aft asymmetry in their interactions. We show here that the resulting antisymmetric part of the "exchange coupling" between a bird and its neighbours, if large enough, destabilizes…

Statistical Mechanics · Physics 2020-05-06 Lokrshi Prawar Dadhichi , Rahul Chajwa , Ananyo Maitra , Sriram Ramaswamy

Numerical models indicate that collective animal behaviour may emerge from simple local rules of interaction among the individuals. However, very little is known about the nature of such interaction, so that models and theories mostly rely…

Populations and Evolution · Quantitative Biology 2009-11-13 M. Ballerini , N. Cabibbo , R. Candelier , A. Cavagna , E. Cisbani , I. Giardina , V. Lecomte , A. Orlandi , G. Parisi , A. Procaccini , M. Viale , V. Zdravkovic

Speed fluctuations of individual birds in natural flocks are moderate, due to the aerodynamic and biomechanical constraints of flight. Yet the spatial correlations of such fluctuations are scale-free, namely they have a range as wide as the…

Understanding collective self-organization in active matter, such as bird flocks and fish schools, remains a grand challenge in physics. Interactions that induce alignment are essential for flocking; however, alignment alone is generally…

Soft Condensed Matter · Physics 2025-07-01 Julian Giraldo-Barreto , Viktor Holubec

Turbulence is omnipresent in the atmosphere and a long-standing scientific conundrum that makes flight complex. This complexity is little understood; surprisingly, when turbulence arises, air vehicles struggle while birds seem to thrive.…

Fluid Dynamics · Physics 2026-01-01 Ariane Gayout , David Lentink

The propulsion of a flapping wing or foil is emblematic of bird flight and fish swimming. Previous studies have identified hallmarks of the propulsive dynamics that have been attributed to unsteady effects such as the formation and shedding…

Fluid Dynamics · Physics 2026-04-07 Olivia Pomerenk , Leif Ristroph

Collective decision-making in biological systems requires all individuals in the group to go through a behavioural change of state. During this transition, the efficiency of information transport is a key factor to prevent cohesion loss and…

This paper investigates stability analysis of flapping flight. Due to time-varying aerodynamic forces, such systems do not display fixed points of equilibrium. The problem is therefore approached via a limit cycle analysis based on Floquet…

We present the first decentralized multi-copter flock that performs stable autonomous outdoor flight with up to 10 flying agents. By decentralized and autonomous we mean that all members navigate themselves based on the dynamic information…

The natural wind environment that volant insects encounter is unsteady and highly complex, posing significant flight control and stability challenges. Unsteady airflows can range from structured chains of discrete vortices shed in the wake…

Biological Physics · Physics 2016-03-03 Sridhar Ravi , Dmitry Kolomenskiy , Thomas Engels , Kai Schneider , Chun Wang , Joern Sesterhenn , Hao Liu

Background: Recent research in animal behaviour has contributed to determine how alignment, turning responses, and changes of speed mediate flocking and schooling interactions in different animal species. Here, we address specifically the…

Quantitative Methods · Quantitative Biology 2018-09-05 Andrea Perna , Guillaume Gregoire , Richard P. Mann

We study the spatial patterns formed by a system of interacting particles where the mobility of any individual is determined by the population crowding at two different spatial scales. In this way we model the behavior of some biological…

Statistical Mechanics · Physics 2015-07-08 Ricardo Martínez-García , Clara Murgui , Emilio Hernández-García , Cristóbal López

Animals that travel together in groups display a variety of fascinating motion patterns thought to be the result of delicate local interactions among group members. Although the most informative way of investigating and interpreting…

Biological Physics · Physics 2010-10-27 Mate Nagy , Zsuzsa Akos , Dora Biro , Tamas Vicsek

Flocks of birds, schools of fish, insects swarms are examples of coordinated motion of a group that arises spontaneously from the action of many individuals. Here, we study flocking behavior from the viewpoint of multi-agent reinforcement…

Physics and Society · Physics 2020-07-08 Mihir Durve , Fernando Peruani , Antonio Celani

Natural flocks (aligned) and swarms (non-aligned) both exhibit features of near-criticality, challenging their treatment as two ends of the same phase transition. We present a model for the aggregation of active individuals, in which their…

Adaptation and Self-Organizing Systems · Physics 2025-11-26 Joao Lizárraga , Marcus de Aguiar

Experiments find coherent information transfer through biological groups on length and time scales distinctly below those on which asymptotically correct hydrodynamic theories apply. We present here a new continuum theory of collective…

The self-organised motion of vast numbers of creatures in a single direction is a spectacular example of emergent order. We recreate this phenomenon using actuated non-living components. We report here that millimetre-sized tapered rods,…

Soft Condensed Matter · Physics 2014-09-23 Nitin Kumar , Harsh Soni , Sriram Ramaswamy , A. K. Sood

We numerically examine a bidisperse system of active and passive particles coupled to a resource substrate. The active particles deplete the resource at a fixed rate and move toward regions with higher resources, while all of the particles…

Soft Condensed Matter · Physics 2024-04-23 L. Varga , A. Libal , C. Reichhardt , C. J. O. Reichhardt

Coordinated collective motion in bird flocks and fish schools inspires algorithms for cohesive swarm robotics. This paper presents a position-based flocking model that achieves persistent velocity alignment without velocity sensing. By…