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Many cell types spontaneously order like nematic liquid crystals, and, as such, they form topological defects. While defects with topological charge $\pm$1/2 are common in cell monolayers, the defects with charge $\pm$1, relevant in the…

Soft Condensed Matter · Physics 2019-12-09 Kirsten D. Endresen , MinSu Kim , Francesca Serra

The role of boundary layers in conventional liquid crystals is commonly subsumed in their anchoring on confining walls. In the classical view, anchoring enslaves the orientational field of the passive material under equilibrium conditions.…

Directing individual motions of many constituents to coherent dynamical state is a fundamental challenge in multiple fields. Here, based on the spherical crystal model, we show that topological defects in particle arrays can be a crucial…

Soft Condensed Matter · Physics 2019-06-11 Zhenwei Yao

Directional media, such as nematic liquid crystals and ferromagnets, are characterized by their topologically stabilized defects in directional order. In nematics, boundary conditions and surface-treated inclusions often create complex…

Soft Condensed Matter · Physics 2015-06-03 Simon Čopar , Slobodan Žumer

Defects are a ubiquitous feature of ordered media. They have certain universal features, independent of the underlying physical system, reflecting their topological origins. While the topological properties of defects are robust, they…

Soft Condensed Matter · Physics 2021-02-24 Chiqun Zhang , Amit Acharya , Alan C. Newell , Shankar C. Venkataramani

Cell layers are often categorized as contractile or extensile active nematics but recent experiments on neural progenitor cells with induced $+1$ topological defects challenge this classification. In a bottom-up approach, we first study a…

Soft Condensed Matter · Physics 2024-09-13 Zihui Zhao , Yisong Yao , He Li , Yongfeng Zhao , Yujia Wang , Hepeng Zhang , Hugues Chat'e , Masaki Sano

We use molecular dynamics to study the ordering of a nematic liquid crystal around a spherical particle or droplet. Homeotropic boundary conditions and strong anchoring create a hedgehog director configuration on the particle surface and in…

Soft Condensed Matter · Physics 2013-03-19 D. Andrienko , G. Germano , M. P. Allen

Understanding the fundamental mechanisms behind plastic instabilities and shear band formation in amorphous media under applied deformation remains a long-standing challenge. Leveraging on the mathematical concept of topology, we revisit…

Soft Condensed Matter · Physics 2025-07-08 Xin Wang , Jin Shang , Yujie Wang , Jie Zhang , Matteo Baggioli

Collective cell migration governs a range of physiological and pathological processes, from tissue morphogenesis to cancer invasion, in which topological defects arise as an inevitable consequence of frequent cellular rearrangement and…

Soft Condensed Matter · Physics 2026-04-01 Jiusi Zhang , Chung Wing Chan , Bo Li , Rui Zhang

When considering flows in biological membranes, they are usually treated as flat, though more often than not, they are curved surfaces, even extremely curved, as in the case of the endoplasmic reticulum. Here, we study the topological…

Fluid Dynamics · Physics 2021-05-27 Rickmoy Samanta , Naomi Oppenheimer

We study the organization of topological defects in a system of nematogens confined to the two-dimensional sphere (S^2). We first perform Monte Carlo simulations of a fluid system of hard rods (spherocylinders) living in the tangent plane…

Soft Condensed Matter · Physics 2009-11-13 Homin Shin , Mark J. Bowick , Xiangjun Xing

Active matter encompasses different nonequilibrium systems in which individual constituents convert energy into non-conservative forces or motion at the microscale. This review provides an elementary introduction to the role of topology in…

Soft Condensed Matter · Physics 2022-06-28 Suraj Shankar , Anton Souslov , Mark J. Bowick , M. Cristina Marchetti , Vincenzo Vitelli

Recent experimental observations have suggested that topological defects can facilitate the creation of sharp features in developing embryos. Whereas these observations echo established knowledge about the interplay between geometry and…

Soft Condensed Matter · Physics 2023-07-25 Ludwig A. Hoffmann , Livio Nicola Carenza , Luca Giomi

The ability to reroute and control flow is vital to the function of venation networks across a wide range of organisms. By modifying individual edges in these networks, either by adjusting edge conductances or creating and destroying edges,…

Soft Condensed Matter · Physics 2021-01-14 Jason W. Rocks , Andrea J. Liu , Eleni Katifori

Efficient nutrient mixing is crucial for the survival of bacterial colonies and other living systems. This raises the question of whether the optimization of mixing through the emergence of active turbulent motion in bacterial swarms played…

Soft Condensed Matter · Physics 2022-09-27 Rodrigo C. V. Coelho , Nuno A. M. Araújo , Margarida M. Telo da Gama

We investigate the dynamics of active nematic liquid crystals on deformable membranes, focusing on the interplay between active stress and anisotropic curvature coupling. Using a minimal model, we simulate the coupled evolution of the…

Soft Condensed Matter · Physics 2026-05-19 Yuki Hirota , Nariya Uchida

We present a hydrodynamic model for a thin spherical shell of active nematic liquid crystal with an arbitrary configuration of defects. The active flows generated by defects in the director lead to the formation of stable vortices,…

Soft Condensed Matter · Physics 2017-11-13 Diana Khoromskaia , Gareth P. Alexander

The effects of an electric field on the flow patterns and defect dynamics of two-dimensional active nematics are numerically investigated. We found that field-induced director reorientation causes anisotropic active turbulence characterized…

Soft Condensed Matter · Physics 2023-07-20 Yutaka Kinoshita , Nariya Uchida

We study how confinement transforms the chaotic dynamics of bulk microtubule-based active nematics into regular spatiotemporal patterns. For weak confinements, multiple continuously nucleating and annihilating topological defects…

Topological defects play a key role in two-dimensional active nematics, and a transient role in two-dimensional active polar fluids. In this paper, we study both the transient and long-time behavior of defects in two-dimensional active…

Soft Condensed Matter · Physics 2020-09-24 Farzan Vafa