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The cytoskeleton is an inhomogeneous network of semi-flexible filaments, which are involved in a wide variety of active biological processes. Although the cytoskeletal filaments can be very stiff and embedded in a dense and cross-linked…

Biological Physics · Physics 2019-05-24 Ines Weber , Cécile Appert-Rolland , Grégory Schehr , Ludger Santen

The actin cytoskeleton is a key component in the machinery of eukaryotic cells, and it selfassembles out of equilibrium into a wide variety of biologically crucial structures. While the molecular mechanisms involved are well characterized,…

Soft Condensed Matter · Physics 2017-01-04 Giulia Foffano , Nicolas Levernier , Martin Lenz

Myosin motor proteins drive vigorous steady-state fluctuations in the actin cytoskeleton of cells. Endogenous embedded semiflexible filaments such as microtubules, or added filaments such as single-walled carbon nanotubes are used as novel…

Biological Physics · Physics 2016-06-21 J. Gladrow , N. Fakhri , F. C. MacKintosh , C. F. Schmidt , C. P. Broedersz

The fracture and severing of polymer chains plays a critical role in the failure of fibrous materials and the regulated turnover of intracellular filaments. Using continuum wormlike chain models, we investigate the fracture of semiflexible…

Statistical Mechanics · Physics 2020-03-05 Alexander Lorenzo , Enrique M. De La Cruz , Elena F. Koslover

The cytoskeleton is an important subsystem of cells that is involved for example in cell division and locomotion. It consists of filaments that are cross-linked by molecular motors that can induce relative sliding between filaments and…

Statistical Mechanics · Physics 2007-08-08 P. K. Mohanty , K. Kruse

We examine the equilibrium fluctuation spectrum of a constituent semiflexible filament segment in a network. The effect of this cross linking is to modify the mechanical boundary conditions at the end of the filament. We consider the effect…

Soft Condensed Matter · Physics 2020-01-22 Jonathan Kernes , Alex J. Levine

The wrapping equilibria of one and two adsorbing cylinders are studied along a semi-flexible filament (polymer) due to the interplay between elastic rigidity and short-range adhesive energy between the cylinder and the filament. We show…

Soft Condensed Matter · Physics 2015-06-05 David S. Dean , Thomas C. Hammant , Ronald R. Horgan , Ali Naji , Rudolf Podgornik

We develop a scaling theory to describe dynamic fluctuations of a semiflexible polymer and find several distinct regimes. We performed simulations to characterize the longitudinal and transverse dynamics; using ensemble averaging for a…

Soft Condensed Matter · Physics 2009-10-31 R. Everaers , F. Julicher , A. Ajdari , A. C. Maggs

Semiflexible slender filaments are ubiquitous in nature and cell biology, including in the cytoskeleton, where reorganization of actin filaments allows the cell to move and divide. Most methods for simulating semiflexible inextensible…

Numerical Analysis · Mathematics 2023-05-03 Ondrej Maxian , Brennan Sprinkle , Aleksandar Donev

While actin bundles are used by living cells for structural fortification, the microscopic origin of the elasticity of bundled networks is not understood. Here, we show that above a critical concentration of the actin binding protein…

Soft Condensed Matter · Physics 2007-08-30 O. Lieleg , M. M. A. E. Claessens , C. Heussinger , E. Frey , A. R. Bausch

We investigate the dynamics of a single semiflexible filament, under the action of a compressing force, using numerical simulations and scaling arguments. The force is applied along the end to end vector at one extremity of the filament,…

Soft Condensed Matter · Physics 2009-11-07 P. Ranjith , P. B. Sunil Kumar

Thermal fluctuations of microtubules (MTs) and other cytoskeletal filaments govern to a great extent the complex rheological properties of the cytoskeleton in eukaryotic cells. In recent years, much effort has been put into capturing the…

Soft Condensed Matter · Physics 2011-10-26 Sander L. Poelert , Harrie H. Weinans , Amir A. Zadpoor

Biological activity gives rise to non-equilibrium fluctuations in the cytoplasm of cells; however, there are few methods to directly measure these fluctuations. Using a reconstituted actin cytoskeleton, we show that the bending dynamics of…

Soft Condensed Matter · Physics 2009-11-13 C. P. Brangwynne , G. H. Koenderink , F. C. MacKintosh , D. A. Weitz

The phase behavior of charged rods in the presence of inter-rod linkers is studied theoretically as a model for the equilibrium behavior underlying the organization of actin filaments by linker proteins in the cytoskeleton. The presence of…

Soft Condensed Matter · Physics 2016-08-31 Itamar Borukhov , Robijn F. Bruinsma , William M. Gelbart , Andrea J. Liu

We construct a coarse-grained model of parallel actin bundles crosslinked by compact, globular bundling proteins, such as fascin and espin, necessary components of filapodial and mechanosensory bundles. Consistent with structural…

Soft Condensed Matter · Physics 2013-05-29 Homin Shin , Gregory M. Grason

Actin is a key component of the cytoskeleton, which plays central roles in cell motility, division, growth, and tensile strength. To enable this wide range of transient mechanical processes and properties, networks of actin filaments…

The nonlinear dynamics of an elastic filament that is forced to rotate at its base is studied by hydrodynamic simulation techniques; coupling between stretch, bend, twist elasticity and thermal fluctuations is included. The…

Soft Condensed Matter · Physics 2008-03-24 Hirofumi Wada , Roland R. Netz

Disordered networks of semiflexible filaments are common support structures in biology. Familiar examples include fibrous matrices in blood clots, bacterial biofilms, and essential components of cells and tissues of plants, animals, and…

Soft Condensed Matter · Physics 2023-09-28 Saamiya Syed , Fred C. MacKintosh , Jordan L. Shivers

We review the physical properties of macromolecular networks, consisting of semiflexible polymers such as actin. We start by giving a theoretical analysis of the conformational statistics and mechanical response of single filaments.…

Soft Condensed Matter · Physics 2008-02-03 Erwin Frey , Klaus Kroy , Jan Wilhelm , Erich Sackmann

Intermediate filaments are the least explored among the large cytoskeletal elements. We show here that they display conformational anomalies in narrow microfluidic channels. Their unusual behavior can be understood as the consequence of a…

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