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A novel two-leaflet description of lipid membranes is proposed. Within its framework, phase separation phenomena in multicomponent biological membranes are analyzed. As we show, interactions between the leaflets tend to suppress macroscopic…

Biological Physics · Physics 2016-02-03 Ramon Reigada , Alexander S. Mikhailov

Reaction-diffusion systems where transition rates exhibit quenched disorder are common in physical and chemical systems. We study pair reactions on a periodic two-dimensional lattice, including continuous deposition and spontaneous…

Disordered Systems and Neural Networks · Physics 2010-11-10 A. Wolff , I. Lohmar , J. Krug , Y. Frank , O. Biham

Membraneless droplets or liquid condensates formed via liquid-liquid phase separation (LLPS) play a pivotal role in cell biology and hold potential for biomedical engineering. While membraneless droplets are often studied in the context of…

Soft Condensed Matter · Physics 2025-11-07 Chenxi Liu , Ding Cao , Siyu Liu , Yilin Wu

Living systems must maintain robust biochemical function despite fluctuations that span a wide range of timescales. Biomolecular condensates formed by liquid-liquid phase separation (LLPS) have been shown to buffer concentration…

Biological Physics · Physics 2025-10-24 Logan de Monchaux-Irons , T-Y Dora Tang , Christoph A. Weber , Thomas C. T. Michaels

Disentangling the mechanistic details of a chemical reaction pathway is a hard problem that often requires a considerable amount of chemical intuition and a component of luck. Experiments struggle in observing short-life metastable…

Chemical Physics · Physics 2019-08-01 Valerio Rizzi , Dan Mendels , Emilia Sicilia , Michele Parrinello

We study analytically kinetics of an elementary bimolecular reaction scheme of the Langmuir-Hinshelwood type taking place on a d-dimensional catalytic substrate. We propose a general approach which takes into account explicitly the…

Statistical Mechanics · Physics 2009-10-31 P. Argyrakis , S. F. Burlatsky , E. Clement , G. Oshanin

Liquid-liquid phase separation (LLPS) is currently of great interest in cell biology. LLPS is an example of what is called an emergent phenomenon -- an idea that comes from condensed-matter physics. Emergent phenomena have the…

Cell Behavior · Quantitative Biology 2021-03-16 Richard P Sear

Every mathematical model describing physical phenomena is an approximation to model reality, hence has its limitations. Depending on characteristic values of the variables in the model, different aspects of the model and, e.g.,…

Analysis of PDEs · Mathematics 2019-12-02 Björn Augner , Dieter Bothe

In a companion paper, we put forth a thermodynamic model for complex formation via a chemical reaction involving multiple macromolecular species, which may subsequently undergo liquid-liquid phase separation and a further transition into a…

Biological Physics · Physics 2024-07-26 Ruoyao Zhang , Sheng Mao , Mikko P. Haataja

The quantitative description of marine systems is constrained by a major issue of scale separation: most marine biochemical processes occur at sub-centimeter scales, while the contribution to the Earth's biogeochemical cycles is expressed…

Atmospheric and Oceanic Physics · Physics 2019-11-14 Francesco Paparella , Marcello Vichi

We demonstrate that the Michaelis-Menten reaction mechanism can be accurately approximated by a linear system when the initial substrate concentration is low. This leads to pseudo-first-order kinetics, simplifying mathematical calculations…

Dynamical Systems · Mathematics 2024-03-11 Justin Eilertsen , Santiago Schnell , Sebastian Walcher

The role of dimensionality (Euclidean versus fractal), spatial extent, boundary effects and system topology on the efficiency of diffusion-reaction processes involving two simultaneously-diffusing reactants is analyzed. We present…

Statistical Mechanics · Physics 2009-11-10 Jonathan L. Bentz , John J. Kozak , E. Abad , G. Nicolis

Proteins can form droplets via liquid-liquid phase separation (LLPS) in cells. Recent experiments demonstrate that LLPS is qualitatively different on two-dimensional (2d) surfaces compared to three-dimensional (3d) solutions. In this paper,…

Quantitative Methods · Quantitative Biology 2024-08-29 Jinyoung Kim , Sean D. Lawley , Jinsu Kim

Many intercalation compounds possess layered structures or inter-penetrating lattices that enable phase separation into three or more stable phases, or "stages," driven by competing intra-layer and inter-layer forces. While these structures…

Chemical Physics · Physics 2017-06-27 Raymond B. Smith , Edwin Khoo , Martin Z. Bazant

We use a standard bead-spring model and molecular dynamics simulations to study the static properties of symmetric linear multiblock copolymer chains and their blocks under poor solvent conditions in a dilute solution from the regime close…

Soft Condensed Matter · Physics 2012-03-05 Panagiotis E. Theodorakis , Nikolaos G. Fytas

The cellular cytoplasm is organized into compartments. Phase separation is a simple manner to create membrane-less compartments in order to confine and localize particles like proteins. In many cases these particles are bound to fluctuating…

Statistical Mechanics · Physics 2020-09-16 G. David , J. -C. Walter , C. P. Broedersz , J. Dorignac , F. Geniet , A. Parmeggiani , N. -O. Walliser , J. Palmeri

In this paper we apply a finite difference lattice Boltzmann model to study the phase separation in a two-dimensional liquid-vapor system. Spurious numerical effects in macroscopic equations are discussed and an appropriate numerical scheme…

Soft Condensed Matter · Physics 2009-11-10 V. Sofonea , A. Lamura , G. Gonnella , A. Cristea

Surfaces serve as highly efficient catalysts for a vast variety of chemical reactions. Typically, such surface reactions involve billions of molecules which diffuse and react over macroscopic areas. Therefore, stochastic fluctuations are…

Statistical Mechanics · Physics 2007-10-12 B. Barzel , O. Biham

The need to use a short time step is a key limit on the speed of molecular dynamics (MD) simulations. Simulations governed by classical potentials are often accelerated by using a multiple-time-step (MTS) integrator that evaluates certain…

Chemical Physics · Physics 2023-10-24 Xiang Fu , Albert Musaelian , Anders Johansson , Tommi Jaakkola , Boris Kozinsky

Motivated by the recent experimental observations on clustering of motor proteins on microtubule filament, we study an open system of two parallel totally asymmetric simple exclusion processes under asymmetric coupling conditions, which…

Statistical Mechanics · Physics 2016-03-23 Arvind Kumar Gupta