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We distinguish a mechanical representation of the world in terms of point masses with positions and momenta and the chemical representation of the world in terms of populations of different individuals, each with intrinsic stochasticity,…

Statistical Mechanics · Physics 2019-05-07 Hong Qian

One of the aims of systems biology is to build multiple layered and multiple scale models of living systems which can efficiently describe phenomena occurring at various level of resolution. Such models should consist of layers of various…

Dynamical Systems · Mathematics 2015-03-03 Jacek Banasiak , Aleksandra Falkiewicz , Proscovia Namayanja

Chemical reaction networks underpin biological and physical phenomena across scales, from microbial interactions to planetary atmosphere dynamics. Bacterial communities exhibit complex competitive interactions for resources, human organs…

Molecular Networks · Quantitative Biology 2026-01-08 Jorge Reyes , Jörn Dunkel

It has long been hypothesized that futile cycles in cellular metabolism are involved in the regulation of biochemical pathways. Following the work of Newsholme and Crabtree, we develop a quantitative theory for this idea based on…

Subcellular Processes · Quantitative Biology 2007-05-23 Hong Qian , Daniel A. Beard

Many mathematical models for biological phenomena, such as the spread of diseases, are based on reaction-diffusion equations for densities of interacting cell populations. We present a consistent derivation of reaction-diffusion equations…

Analysis of PDEs · Mathematics 2026-02-23 Marzia Bisi , Davide Cusseddu , Ana Jacinta Soares , Romina Travaglini

It is shown that the dual to the linear programming problem that arises in constraint-based models of metabolism can be given a thermodynamic interpretation in which the shadow prices are chemical potential analogues, and the objective is…

Subcellular Processes · Quantitative Biology 2009-11-13 Patrick B. Warren , Janette L. Jones

As the quantification of metabolism, nonequilibrium steady states play a central role in living matter, but are beyond the purview of equilibrium statistical mechanics. Here we develop a fermionic theory of nonequilibrium steady states in…

Statistical Mechanics · Physics 2024-04-24 Eric De Giuli , Masanari Shimada

Biological systems perform an astonishing array of dynamical processes -- including development and repair, regulation, behavior and motor control, sensing and signaling, and adaptation, among others. Powered by the transduction of stored…

Soft Condensed Matter · Physics 2025-09-04 James Clarke , Jake McGrath , Colin Johnson , José Alvarado

Living systems are forced away from thermodynamic equilibrium by exchange of mass and energy with their environment. In order to model a biochemical reaction network in a non-equilibrium state one requires a mathematical formulation to…

Molecular Networks · Quantitative Biology 2012-02-21 Ronan M. T. Fleming , Ines Thiele

A major goal of modern computational biology is to simulate the collective behaviour of large cell populations starting from the intricate web of molecular interactions occurring at the microscopic level. In this paper we describe a…

Biological Physics · Physics 2007-05-23 Roberto Chignola , Edoardo Milotti

Nonlinear density response theory is revisited focusing on the harmonically perturbed finite temperature uniform electron gas. Within the non-interacting limit, brute force quantum kinetic theory calculations for the quadratic, cubic,…

Quantum Gases · Physics 2023-05-23 Panagiotis Tolias , Tobias Dornheim , Zhandos A. Moldabekov , Jan Vorberger

We study the multi-species replicator model with linear fitness and random fitness matrices of various classes. By means of numerical resolution of the replicator equations, we determine the survival probability of a species in terms of its…

Adaptation and Self-Organizing Systems · Physics 2007-05-23 Sebastian Bouzat , Damian Zanette

We consider the electrons of a molecule in the adiabatic time-dependent density functional theory approximation. We establish the well-posedness of the time evolution and its linear response close to a non-degenerate ground state, and prove…

Analysis of PDEs · Mathematics 2024-09-19 Mi-Song Dupuy , Eloïse Letournel , Antoine Levitt

One of the major research questions regarding human microbiome studies is the feasibility of designing interventions that modulate the composition of the microbiome to promote health and cure disease. This requires extensive understanding…

Methodology · Statistics 2021-11-18 Matthew D. Koslovsky , Kristi L. Hoffman , Carrie R. Daniel , Marina Vannucci

Controlling complex reaction networks is a fundamental challenge in the fields of physics, biology, and systems engineering. Here, we prove a general principle for catalytic reaction systems with kinetics where the reaction order and the…

Biological Physics · Physics 2026-02-26 Yusuke Himeoka , Shuhei A. Horiguchi , Naoto Shiraishi , Fangzhou Xiao , Tetsuya J. Kobayashi

Basing on the theory of Feynman's influence functional and its hierarchical equations of motion, we develop a linear response theory for quantum open systems. Our theory provides an effective way to calculate dynamical observables of a…

Mesoscale and Nanoscale Physics · Physics 2011-08-31 J. H. Wei , YiJing Yan

We introduce a comprehensive analytical benchmark, relying on Fokker-Planck formalism, to study microbial dynamics in presence of both biotic and abiotic forces. In equilibrium, we observe a balance between the two kinds of forces, leading…

Populations and Evolution · Quantitative Biology 2024-10-17 José Camacho-Mateu , Aniello Lampo , Saúl Ares , José A. Cuesta

System-level properties of metabolic networks may be the direct product of natural selection or arise as a by-product of selection on other properties. Here we study the effect of direct selective pressure for growth or viability in…

Molecular Networks · Quantitative Biology 2015-08-03 Marco Bardoscia , Matteo Marsili , Areejit Samal

The Green-Kubo relation, the Einstein relation, and the fluctuation-response relation are representative universal relations among measurable quantities that are valid in the linear response regime. We provide pedagogical proofs of these…

Statistical Mechanics · Physics 2009-11-11 Kumiko Hayashi , Shin-ichi Sasa

Intracellular protein patterns govern essential cellular functions by dynamically redistributing proteins between membrane-bound and cytosolic states, conserving their total numbers. This review presents a theoretical framework for…

Biological Physics · Physics 2025-12-16 Erwin Frey , Henrik Weyer