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Swimming microorganisms often self propel in fluids with complex rheology. While past theoretical work indicates that fluid viscoelasticity should hinder their locomotion, recent experiments on waving swimmers suggest a possible…

Biological Physics · Physics 2014-11-25 Emily E. Riley , Eric Lauga

Microorganisms are rarely found in Nature swimming freely in an unbounded fluid. Instead, they typically encounter other organisms, hard walls, or deformable boundaries such as free interfaces or membranes. Hydrodynamic interactions between…

Fluid Dynamics · Physics 2013-10-21 Marcelo A. Dias , Thomas R. Powers

We study the stochastic force dynamics of a model microswimmer (Chlamydomonas reinhardtii), using a combined experimental, theoretical, and numerical approach. While swimming dynamics have been extensively studied using hydrodynamic…

We consider the dynamics of micro-sized, asymmetrically-coated thermoresponsive hydrogel ribbons (microgels) under periodic heating and cooling in the confined space between two planar surfaces. As the result of the temperature changes, the…

Soft Condensed Matter · Physics 2022-03-23 Ivan Tanasijević , Oliver Jung , Lyndon Koens , Ahmed Mourran , Eric Lauga

A theory is presented for wave-driven propulsion of floating bodies driven into oscillation at the fluid interface. By coupling the equations of motion of the body to a quasi-potential flow model of the fluid, we derive expressions for the…

Fluid Dynamics · Physics 2023-10-20 GP Benham , O Devauchelle , SJ Thomson

The physical mechanism of microbial motion near solid-liquid interfaces is crucial for understanding various biological phenomena and developing ecological applications. However, limited works have been conducted on the swimming behavior of…

Biological Physics · Physics 2024-11-14 Chunhe Li , Hongyi Bian , Yateng Qiao , Jin Zhu , Zijie Qu

Much current study of legged locomotion has rightly focused on foot traction forces, including on granular media. Future legged millirobots will need to go through terrain, such as brush or other vegetation, where the body contact forces…

Robotics · Computer Science 2019-04-22 Can Koc , Cem Koc , Brian Su , Carlos Casarez , Ronald Fearing

The physical and bio-mechanical principles that govern undulatory movement on wet surfaces have important applications in physiology, physics, and engineering. The nematode {\it C. elegans}, with its highly stereotypical and functionally…

Fluid Dynamics · Physics 2015-06-03 Xiao N. Shen , J. Sznitman , P. Krajacic , T. Lamitina , P. E. Arratia

Many microorganisms swim by performing larger non-reciprocal shape deformations that are initiated locally by molecular motors. However, it remains unclear how decentralized shape control determines the movement of the entire organism.…

Biological Physics · Physics 2025-05-14 Benedikt Hartl , Michael Levin , Andreas Zöttl

A number of swimming microorganisms such as ciliates ($\textit{Opalina}$) and multicellular colonies of flagellates ($\textit{Volvox}$) are approximately spherical in shape and swim using beating arrays of cilia or short flagella covering…

Fluid Dynamics · Physics 2014-02-06 On Shun Pak , Eric Lauga

Microorganisms often encounter strong confinement and complex hydrodynamic flows while navigating their habitats. Combining finite-element methods and stochastic simulations, we study the interplay of active transport and heterogeneous…

Soft Condensed Matter · Physics 2025-11-05 Pallabi Das , Mirko Residori , Axel Voigt , Suvendu Mandal , Christina Kurzthaler

Many microswimmers are able to swim through viscous fluids by employing periodic non-reciprocal deformations of their appendages. Here we use a simple microswimmer model inspired by swimming biflagellates which consists of a spherical cell…

Soft Condensed Matter · Physics 2025-08-22 Sridhar Bulusu , Andreas Zöttl

The complex formations exhibited by schooling fish have long been the object of fascination for biologists and physicists. However, the physical and sensory mechanisms leading to organized collective behavior remain elusive. On the physical…

Fluid Dynamics · Physics 2025-07-09 Anand U. Oza , Eva Kanso , Michael J. Shelley

The motion of microswimmers in complex flows is ruled by the interplay between swimmer propulsion and the dynamics induced by the fluid velocity field. Here we study the motion of a chiral microswimmer whose propulsion is provided by the…

Fluid Dynamics · Physics 2021-06-08 Ji Zhang , Mauro Chinappi , Luca Biferale

Locomotion on small scales is dominated by the effects of viscous forces and, as a result, is subject to strong physical and mathematical constraints. Following Purcell's statement of the scallop theorem which delimitates the types of…

Biological Physics · Physics 2011-08-30 Eric Lauga

We present and analyze a theoretical model for the dynamics and interactions of "capillary surfers," which are millimetric objects that self-propel while floating at the interface of a vibrating fluid bath. In our companion paper [1], we…

Fluid Dynamics · Physics 2024-11-25 Anand U. Oza , Giuseppe Pucci , Ian Ho , Daniel M. Harris

We investigate the hydrodynamic interactions between microorganisms swimming at low Reynolds number. By considering simple model swimmers, and combining analytic and numerical approaches, we investigate the time-averaged flow field around a…

Soft Condensed Matter · Physics 2007-05-25 C. M. Pooley , G. P. Alexander , J. M. Yeomans

An approximation to the added mass matrix of an assembly of spheres is constructed on the basis of potential flow theory for situations where one sphere is much larger than the others. In the approximation the flow potential near a small…

Fluid Dynamics · Physics 2017-03-08 B. U. Felderhof

Motivated by recent experiments of motile bacteria crossing liquid-liquid interfaces of isotropic- nematic coexistence (Cheon et al., Soft Matter 20: 7313-7320, 2024), we study the dynamics of prolate microswimmers traversing clean…

Soft Condensed Matter · Physics 2025-08-01 Rishish Mishra , Harish Pothukuchi , Harinadha Gidituri , Juho Lintuvuori

Flagella and cilia are cellular appendages that inherit essential functions of microbial life including sensing and navigating the environment. In order to propel a swimming microorganism they displace the surrounding fluid by means of…