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At the millimeter scale and in the intermediate Reynolds number (Re) regime, the midge and mosquito larvae can reach swimming speeds of more than one body length per cycle performing a "figure-of-8" gait, in which their elongated bodies…

Fluid Dynamics · Physics 2023-03-29 Bowen Jin , Chengfeng Pan , Neng Xia , Jialei Song , Haoxiang Luo , Li Zhang , Yang Ding

Swimming at a micrometer scale demands particular strategies. Indeed when inertia is negligible as compared to viscous forces (i.e. Reynolds number $Re$ is lower than unity), hydrodynamics equations are reversible in time. To achieve…

Soft Condensed Matter · Physics 2011-02-17 Michaël Garcia , Stefano Berti , Philippe Peyla , Salima Rafaï

Self-propelling organisms locomote via generation of patterns of self-deformation. Despite the diversity of body plans, internal actuation schemes and environments in limbless vertebrates and invertebrates, such organisms often use similar…

A body immersed in a supersaturated fluid like carbonated water can accumulate a dynamic field of bubbles upon its surface. If the body is mobile, the attached bubbles can lift it upward against gravity, but a fluid-air interface can clean…

Fluid Dynamics · Physics 2024-05-10 Saverio E. Spagnolie , Samuel Christianson , Carsen Grote

In this paper we study the locomotion of a shape-changing body swimming in a two-dimensional perfect fluid of infinite extent. The shape-changes are prescribed as functions of time satisfying constraints ensuring that they result from the…

Optimization and Control · Mathematics 2009-10-29 Thomas Chambrion , Alexandre Munnier

In the study of microscopic flows, self-propulsion has been particularly topical in recent years, with the rise of miniature artificial swimmers as a new tool for flow control, low Reynolds number mixing, micromanipulation or even drug…

We present in this fluid dynamics video a novel experimental setup with self-propelled swimmers on a free surface. The swimmers, modeled as flexible thin filaments, are subjected to external electromagnetic forcing driving a propagating…

Fluid Dynamics · Physics 2014-11-04 Sophie Ramananarivo , Benjamin Thiria , Ramiro Godoy-Diana

Many small organisms self-propel in viscous fluids using travelling wave-like deformation of their bodies or appendages. Examples include small nematodes moving through soil using whole-body undulations or spermatozoa swimming through mucus…

Biological Physics · Physics 2015-07-02 Emily E. Riley , Eric Lauga

The properties of biological microswimmers are to a large extent determined by fluid-mediated interactions, which govern their propulsion, perception of their surrounding, and the steering of their motion for feeding or in pursuit.…

Soft Condensed Matter · Physics 2024-06-04 Segun Goh , Roland G. Winkler , Gerhard Gompper

The locomotion of microorganisms and spermatozoa in complex viscoelastic fluids is of critical importance in many biological processes such as fertilization, infection, and biofilm formation. Depending on their propulsion mechanisms,…

Soft Condensed Matter · Physics 2021-09-14 Gaojin Li , Eric Lauga , Arezoo M. Ardekani

Flight on the two-dimensional air-water interface, with body weight supported by surface tension, is a unique locomotion strategy well adapted for the environmental niche on the surface of water. Although previously described in…

Fluid Dynamics · Physics 2016-01-11 Haripriya Mukundarajan , Thibaut C Bardon , Dong Hyun Kim , Manu Prakash

Undulatory locomotion is a means of self-propulsion that relies on the generation and propagation of waves along a body. As a mode of locomotion it is primitive and relatively simple, yet can be remarkably robust. No wonder then, that it is…

Biological Physics · Physics 2009-08-20 Netta Cohen , Jordan H. Boyle

Many microorganisms and artificial microswimmers use helical appendages in order to generate locomotion. Though often rotated so as to produce thrust, some species of bacteria such Spiroplasma, Rhodobacter sphaeroides and Spirochetes induce…

Biological Physics · Physics 2018-10-23 Lyndon Koens , Hang Zhang , Martin Moeller , Ahmed Mourran , Eric Lauga

Microorganisms such as bacteria often swim in fluid environments that cannot be classified as Newtonian. Many biological fluids contain polymers or other heterogeneities which may yield complex rheology. For a given set of boundary…

Fluid Dynamics · Physics 2015-06-30 Gwynn Elfring , Eric Lauga

Locomotion by shape changes (spermatozoon swimming, snake slithering, bird flapping) or gas expulsion (rocket firing) is assumed to require environmental interaction, due to conservation of momentum. As first noted in (Wisdom, 2003) and…

Self-propelled particles can exhibit surprising non-equilibrium behaviors, and how they interact with obstacles or boundaries remains an important open problem. Here we show that chemically propelled micro-rods can be captured, with little…

Soft Condensed Matter · Physics 2014-02-21 Daisuke Takagi , Jeremie Palacci , Adam B. Braunschweig , Michael J. Shelley , Jun Zhang

Centipedes exhibit great maneuverability in diverse environments due to their many legs and body-driven control. By leveraging similar morphologies and control strategies, their robotic counterparts also demonstrate effective terrestrial…

Robotics · Computer Science 2025-06-04 Esteban Flores , Baxi Chong , Daniel Soto , Daniel I. Goldman

Hovering insects are limited by their physiology and need to rotate their wings at the end of each back and forth motion to keep the wing's leading edge ahead of its trailing edge. The wing rotation at the end of each half-stroke pushes the…

Fluid Dynamics · Physics 2022-10-28 Swathi Krishna , Alexander Gehrke , Karen Mulleners

Several micro-organisms, such as bacteria, algae, or spermatozoa, use flagella or cilia to swim in a fluid, while many other micro-organisms instead use ample shape deformation, described as amoeboid, to propel themselves by either crawling…

Biological Physics · Physics 2016-09-19 Hao Wu , A. Farutin , W. -F. Hu , M. Thiébaud , S. Rafaï , P. Peyla , M. -C. Lai , C. Misbah

The time dynamics of flagellar and ciliary beating is often neglected in theories of microswimmers, with the most common models prescribing a time-constant actuation of the surrounding fluid. By explicitly introducing a metachronal wave,…

Soft Condensed Matter · Physics 2026-01-19 G. C. Antunes , H. Stark