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Turing patterns in reaction-diffusion (RD) systems have classically been studied only in RD systems which do not explicitly depend on independent variables such as space. In practise, many systems for which Turing patterning is important…

Analysis of PDEs · Mathematics 2023-01-23 Jacob C. Vandenberg , Mark B. Flegg

In this work, we present and analyze a general framework for vegetation dynamics in arid and semi-arid ecosystems in which non-local interactions are purely competitive. The generality of the formulation enables a systematic search for…

Pattern Formation and Solitons · Physics 2026-04-21 Jelle van der Voort , Ricardo Martinez-Garcia , Arjen Doelman

Reaction-diffusion systems with time-delay defined on complex networks have been studied in the framework of the emergence of Turing instabilities. The use of the Lambert W-function allowed us get explicit analytic conditions for the onset…

Statistical Mechanics · Physics 2016-08-03 Julien Petit , Malbor Asllani , Duccio Fanelli , Ben Lauwens , Timoteo Carletti

Reaction diffusion systems are often used to study pattern formation in biological systems. However, most methods for understanding their behavior are challenging and can rarely be applied to complex systems common in biological…

Analysis of PDEs · Mathematics 2013-05-24 William R. Holmes

Pattern formation often occurs in confined systems, yet how boundaries shape patterning dynamics is unclear. We develop techniques to analyze confinement effects in nonlocal advection-diffusion equations, which generically capture the…

Pattern Formation and Solitons · Physics 2025-09-11 Jan Rombouts , Michael L Zhao , Alexander Aulehla , Anna Erzberger

Forecasting system behaviour near and across bifurcations is crucial for identifying potential shifts in dynamical systems. While machine learning has recently been used to learn critical transitions and bifurcation structures from data,…

Machine Learning · Computer Science 2025-11-14 Eva van Tegelen , George van Voorn , Ioannis Athanasiadis , Peter van Heijster

The formation of protein patterns inside cells is generically described by reaction-diffusion models. The study of such systems goes back to Turing, who showed how patterns can emerge from a homogenous steady state when two reactive…

Biological Physics · Physics 2020-02-26 Manon C. Wigbers , Fridtjof Brauns , Tobias Hermann , Erwin Frey

Turing patterns are stationary, wave-like structures that emerge from the nonequilibrium assembly of reactive and diffusive components. While they are foundational in biophysics, their classical formulation relies on a single characteristic…

Soft Condensed Matter · Physics 2026-01-30 Siamak Mirfendereski , Ankur Gupta

Turing instabilities for a two species reaction-diffusion systems is studied under anisotropic diffusion. More specifically, the diffusion constants which characterize the ability of the species to relocate in space are direction sensitive.…

Statistical Mechanics · Physics 2015-09-30 Daniel M. Busiello , Gwendoline Planchon , Malbor Asllani , Timoteo Carletti , Duccio Fanelli

The problem of pattern formation in a generic two species reaction--diffusion model is studied, under the hypothesis that only one species can diffuse. For such a system, the classical Turing instability cannot take place. At variance, by…

Statistical Mechanics · Physics 2013-09-16 Laura Cantini , Claudia Cianci , Duccio Fanelli , Emma Massi , Luigi Barletti

We investigate Turing instability and pattern formation in two-dimensional domains for two reaction-diffusion models, obtained as diffusive limits of kinetic equations for mixtures of monatomic and polyatomic gases. The first model is of…

Mathematical Physics · Physics 2026-02-23 Stefano Boccelli , Giorgio Martalò , Romina Travaglini

Turing instability in activator-inhibitor systems provides a paradigm of nonequilibrium pattern formation; it has been extensively investigated for biological and chemical processes. Turing pattern formation should furthermore be possible…

Adaptation and Self-Organizing Systems · Physics 2010-05-13 Hiroya Nakao , Alexander S. Mikhailov

Models of diffusion driven pattern formation that rely on the Turing mechanism are utilized in many areas of science. However, many such models suffer from the defect of requiring fine tuning of parameters or an unrealistic separation of…

Other Quantitative Biology · Quantitative Biology 2015-03-17 Thomas Butler , Nigel Goldenfeld

Pattern formation from homogeneity is well-studied, but less is known concerning symmetry-breaking instabilities in heterogeneous media. It is nontrivial to separate observed spatial patterning due to inherent spatial heterogeneity from…

Pattern Formation and Solitons · Physics 2019-12-10 Andrew L. Krause , Václav Klika , Thomas E. Woolley , Eamonn A. Gaffney

Random walks represent an important tool for probing the structural and dynamical properties of networks and modeling transport and diffusion processes on networks. However, when individuals' movement becomes dictated by more complicated…

Pattern Formation and Solitons · Physics 2022-11-24 Per Sebastian Skardal

In this work we investigate the effect of density dependent nonlinear diffusion on pattern formation in the Brusselator system. Through linear stability analysis of the basic solution we determine the Turing and the oscillatory instability…

Mathematical Physics · Physics 2015-06-17 G. Gambino , M. C. Lombardo , M. Sammartino , V. Sciacca

Reaction-diffusion processes across layered media arise in several scientific domains such as pattern-forming E. coli on agar substrates, epidermal-mesenchymal coupling in development, and symmetry-breaking in cell polarisation. We develop…

Pattern Formation and Solitons · Physics 2020-09-18 Andrew L. Krause , Václav Klika , Jacob Halatek , Paul K. Grant , Thomas E. Woolley , Neil Dalchau , Eamonn A. Gaffney

Logistic growth process with nonlocal interactions is considered in one dimension. Spontaneous breakdown of translational invariance is shown to take place at some parameter region, and the bifurcation regime is identified for short and…

Soft Condensed Matter · Physics 2009-11-10 Nadav M. Shnerb

Symmetry-breaking instabilities play an important role in understanding the mechanisms underlying the diversity of patterns observed in nature, such as in Turing's reaction--diffusion theory, which connects cellular signalling and transport…

Pattern Formation and Solitons · Physics 2023-12-25 Andrew L. Krause , Eamonn A. Gaffney , Thomas Jun Jewell , Václav Klika , Benjamin J. Walker

Confirming Turing's theory of morphogens in developmental processes is challenging, and synthetic biology has opened new avenues for testing Turing's predictions. Synthetic mammalian pattern formation has been recently achieved through a…

Pattern Formation and Solitons · Physics 2025-09-22 Mohamed Amine Ouchdiri , Saad Benjelloun , Adnane Saoud , Irene Otero-Muras