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Mixing in viscous fluids is challenging, but chaotic advection in principle allows efficient mixing. In the best possible scenario,the decay rate of the concentration profile of a passive scalar should be exponential in time. In practice,…

Fluid Dynamics · Physics 2013-09-24 Jean-Luc Thiffeault , Emmanuelle Gouillart , Olivier Dauchot

A major challenge in the study of active systems is to harness their non-equilibrium dynamics into useful work. We address this by showing how to design colloids with controllable spontaneous propulsion or rotation when immersed in active…

Soft Condensed Matter · Physics 2023-07-12 Alexander J. H. Houston , Gareth P. Alexander

Monolayers of growing bacteria, confined within channel geometries, exhibit self-organization into a highly aligned laminar state along the axis of the channel. Although this phenomenon has been observed in experiments and simulations under…

Soft Condensed Matter · Physics 2024-11-04 Blake Langeslay , Will Fahy , Gabriel Juarez

Bacterial growth and division generally occur by the process known as binary fission, in which the cells grow polarly until they divide into two daughter cells. Although this process is affected by factors that introduce stochastic…

Soft Condensed Matter · Physics 2022-10-05 Andrés Delgado-Campos , Alejandro Cuetos

The ability of many living systems to actively self-propel underlies critical biomedical, environmental, and industrial processes. While such active transport is well-studied in uniform settings, environmental complexities such as geometric…

Biological Physics · Physics 2021-08-23 Alejandro Martínez-Calvo , Carolina Trenado-Yuste , Sujit S. Datta

Non-equilibrium dynamics of topological defects can be used as a fundamental propulsion mechanism in microscopic active matter. Here, we demonstrate swimming of topological defect-propelled colloidal particles in (passive) nematic fluids…

Soft Condensed Matter · Physics 2021-09-30 Tianyi Yao , Žiga Kos , Yimin Luo , Edward B. Steager , Miha Ravnik , Kathleen J. Stebe

Cell migration is fundamental to development, tissue organization, immune response, and disease progression. Amoeboid motility is distinguished by rapid motion and strongly fluctuating cell shapes, reflecting the intrinsically nonlinear…

Soft Condensed Matter · Physics 2026-05-19 Winfried Schmidt , Chaouqi Misbah , Alexander Farutin

Active nematics are materials composed of mobile, elongated particles that can transform energy from the environment into a mechanical motion. Current experimental realizations of the active nematics are of biological origin and include…

Soft Condensed Matter · Physics 2024-06-05 Andrey Sokolov , Jaideep Katuri , Juan J. de Pablo , Alexey Snezhko

Microbial interactions regulate their spread and survival in competitive environments. It is not clear if the physical parameters of the environment regulate the outcome of these interactions. In this work, we show that the opportunistic…

Cell Behavior · Quantitative Biology 2023-09-12 Divakar Badal , Aloke Kumar , Varsha Singh , Danny Raj M

Cultured stem cells have become a standard platform not only for regenerative medicine and developmental biology but also for biophysical studies. Yet, the characterization of cultured stem cells at the level of morphology and macroscopic…

Soft Condensed Matter · Physics 2022-06-23 Kyogo Kawaguchi , Ryoichiro Kageyama , Masaki Sano

How can a collection of motile cells, each generating contractile nematic stresses in isolation, become an extensile nematic at the tissue-level? Understanding this seemingly contradictory experimental observation, which occurs irrespective…

Soft Condensed Matter · Physics 2022-02-21 Andrew Killeen , Thibault Bertrand , Chiu Fan Lee

Many mature bacterial colonies and biofilms are complex three-dimensional (3D) structures. One key step in their developmental program is a transition from a two-dimensional (2D) monolayer into a 3D architecture. Despite the importance of…

Soft Condensed Matter · Physics 2020-03-13 Sho C. Takatori , Kranthi K. Mandadapu

Active swarms, consisting of individual agents which consume energy to move or produce work, are known to generate a diverse range of collective behaviors. Many examples of active swarms are biological in nature (e.g., fish shoals and bird…

Soft Condensed Matter · Physics 2019-02-06 Vladimir Khodygo , Martin T. Swain , Adil Mughal

Coupling between flow and orientation is a central issue in understanding the collective dynamics of active biofilaments and cells. Active stresses generated by motor activity destroy (quasi-)long-range orientational order and induce…

Soft Condensed Matter · Physics 2025-06-04 Yutaka Kinoshita , Nariya Uchida

Biofilms inhabit a range of environments, such as dental plaques or soil micropores, often characterized by intricate, non-even surfaces. However, the impact of surface irregularities on the population dynamics of biofilms remains elusive…

Populations and Evolution · Quantitative Biology 2023-08-31 Witold Postek , Klaudia Staskiewicz , Elin Lilja , Bartlomiej Waclaw

The interplay between active matter and its environment is central to understanding emergent behavior in biological and synthetic systems. Here, we show that coupling active nematic flows to small-amplitude deformations of a compliant…

Soft Condensed Matter · Physics 2026-01-19 Varun Venkatesh , Amin Doostmohammadi

Variations of transport efficiency in structured environments between distinct individuals in actively self-propelled systems is both hard to study and poorly understood. Here, we study the transport of a non-tumbling {\ecoli} strain, an…

Soft Condensed Matter · Physics 2023-08-28 Pooja Chopra , David Quint , Ajay Gopinathan , Bin Liu

Collective dynamics in proliferating anisotropic particle systems arise from an interplay between growth, division, and mechanical interactions, often mediated by particle shape. In classical models of prolate, rod-like growth, flow-induced…

Soft Condensed Matter · Physics 2025-12-02 Jonas Isensee , Lukas Hupe , Philip Bittihn

A wide range of living and artificial active matter exists in close contact with substrates and under strong confinement, where in addition to dipolar active stresses, quadrupolar active stresses can become important. Here, we numerically…

Soft Condensed Matter · Physics 2024-04-23 Salik A. Sultan , Mehrana R. Nejad , Amin Doostmohammadi

Mixtures of active and passive particles are predicted to exhibit a variety of nonequilibrium phases. Here we report a dynamic clustering phase in mixtures of colloids and motile bacteria. We show that colloidal clustering results from a…

Soft Condensed Matter · Physics 2022-05-25 Shreyas Gokhale , Junang Li , Alexandre Solon , Jeff Gore , Nikta Fakhri