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Vegetation in semi-arid environments self-organizes into striking spatial patterns -- bands, spots, labyrinths, and gaps -- with characteristic wavelengths on the order of tens to hundreds of meters. Existing reaction-diffusion models…

Pattern Formation and Solitons · Physics 2026-04-06 Chad M. Topaz

Lotus leaves floating on water usually experience short-wavelength edge wrinkling that decays toward the center, while the leaves growing above water normally morph into a global bending cone shape with long rippled waves near the edge.…

Soft Condensed Matter · Physics 2020-01-28 Fan Xu , Chenbo Fu , Yifan Yang

We provide a derivation of the Foppl-von Karman equations for the shape of and stresses in an elastic plate with residual strains. These might arise from a range of causes: inhomogeneous growth, plastic deformation, swelling or shrinkage…

Analysis of PDEs · Mathematics 2015-05-18 Marta Lewicka , L. Mahadevan , Reza Pakzad

Growth in confined spaces can drive cellular populations through a jamming transition from a fluid-like state to a solid-like state. Experiments have found that jammed budding yeast populations can build up extreme compressive pressures…

Soft Condensed Matter · Physics 2019-05-29 Pawel Gniewek , Carl F Schreck , Oskar Hallatschek

The paper introduces a PDE model for the growth of a tree stem or a vine. The equations describe the elongation due to cell growth, and the response to gravity and to external obstacles. An additional term accounts for the tendency of a…

Optimization and Control · Mathematics 2020-11-10 Alberto Bressan , Michele Palladino , Wen Shen

Plant morphogenesis relies on dynamic growth deformations at the cell and tissue scales driven by osmotic fluxes. A mechanistic understanding of this phenomenon demands a physical framework that integrates cell imbibition, tissue mechanics,…

Biological Physics · Physics 2025-12-08 Hadrien Oliveri , Christophe Godin , Ibrahim Cheddadi

We model the formation and evolution of wrinkles in a floating elastic sheet under uniaxial compression. This is a canonical setup in the study of wrinkling, and whilst its static equilibrium configuration is well characterised, its…

Fluid Dynamics · Physics 2025-10-31 Daniel J. Netherwood , Ben S. Humphries , Connor Robbins , Doireann O'Kiely

Feedback loops are essential for regulating cell proliferation and maintaining the delicate balance between cell division and cell death. Thanks to the exact solution of a few simple models of cell growth it is by now clear that stochastic…

Populations and Evolution · Quantitative Biology 2023-03-07 Antonio Francesco Zirattu , Marta Biondo , Matteo Osella , Michele Caselle

We investigate how thin sheets of arbitrary shapes morph under the isotropic in-plane expansion of their top surface, which may represent several stimuli such as nonuniform heating, local swelling and differential growth. Inspired by…

Soft Condensed Matter · Physics 2015-11-02 Matteo Pezzulla , Gabriel P. Smith , Paola Nardinocchi , Douglas P. Holmes

Plants are a paradigm for active shape control in response to stimuli. For instance, it is well-known that a tilted plant will eventually straighten vertically, demonstrating the influence of both an external stimulus, gravity, and an…

Biological Physics · Physics 2025-09-09 Hadrien Oliveri , Derek E. Moulton , Heather A. Harrington , Alain Goriely

How much energy does it take to stamp a thin elastic shell flat? Motivated by recent experiments on the wrinkling patterns of floating shells, we develop a rigorous method via $\Gamma$-convergence for answering this question to leading…

Analysis of PDEs · Mathematics 2021-02-17 Ian Tobasco

Understanding how growth induces form is a longstanding biological question. Many studies concentrated on the shapes of plant cells, fungi or bacteria. Some others have shown the importance of the mechanical properties of bacterial walls…

Biological Physics · Physics 2007-05-23 Arezki Boudaoud

We solve several problems that involve imposing metrics on surfaces. The problem of a strip with a linear metric gradient is formulated in terms of a Lagrangean similar to those used for spin systems. We are able to show that the low energy…

Soft Condensed Matter · Physics 2007-05-23 M. Marder , N. Papanicolaou

One of the most important branches of nonlinear control theory is the so-called sliding-mode. Its aim is the design of a (nonlinear) feedback law that brings and maintains the state trajectory of a dynamic system on a given sliding surface.…

Systems and Control · Electrical Eng. & Systems 2021-09-14 Mauro Bisiacco , Gianluigi Pillonetto

Tissue growth underpins a wide array of biological and developmental processes, and numerical modeling of growing systems has been shown to be a useful tool for understanding these processes. However, the phenomena that can be captured are…

Soft Condensed Matter · Physics 2023-11-08 Andrew Killeen , Benjamin Partridge , Thibault Bertrand , Chiu Fan Lee

Proliferating cell collectives often develop an active growth layer near their boundary that regulates expansion and morphology, as observed in systems ranging from bacterial biofilms to epithelial tissues and tumor spheroids. While such…

Soft Condensed Matter · Physics 2026-04-27 Fidel Álvarez-Murphy , Ignacio Medina , Néstor Sepúlveda , Gustavo Düring

Randomly crumpled sheets have shape memory. In order to understand the basis of this form of memory, we simulate triangular lattices of springs whose lengths are altered to create a topography with multiple potential energy minima. We then…

Soft Condensed Matter · Physics 2015-11-18 Naomi Oppenheimer , Thomas A. Witten

A variational framework is introduced to describe how a surface bends when it is subject to local constraints on its geometry. This framework is applied to describe the patterns of a folded sheet of paper. The unstretchability of paper…

Other Condensed Matter · Physics 2008-01-24 Jemal Guven , Martin Michael Mueller

Feedback Alignment (FA) methods are biologically inspired local learning rules for training neural networks with reduced communication between layers. While FA has potential applications in distributed and privacy-aware ML, limitations in…

Machine Learning · Computer Science 2024-06-05 Zachary Robertson , Oluwasanmi Koyejo

We consider the statistics of density number of filaments for the propagation of a laser beam subjected to multiple filamentation in a closed area with reflecting boundaries. Dissipation arrests the catastrophic collapse of filaments,…

Optics · Physics 2012-08-21 A. D. Bulygin