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The persistent dynamics in systems out of equilibrium, particularly those characterized by annihilation and creation of topological defects, is known to involve complicated spatiotemporal processes and is deemed difficult to control. Here…

Soft Condensed Matter · Physics 2022-11-22 Zhi-Feng Huang , Hartmut Löwen , Axel Voigt

The hydrodynamic theory of active nematics has been often used to describe the spatio-temporal dynamics of cell flows and motile topological defects within soft confluent tissues. Those theories, however, often rely on the assumption that…

Soft Condensed Matter · Physics 2023-08-15 Ioannis Hadjifrangiskou , Liam J. Ruske , Julia M. Yeomans

Active nematic models explain the topological defects and flow patterns observed in epithelial tissues, but the nature of active stress-whether it is extensile or contractile, a key parameter of the theory-is not well established…

Soft Condensed Matter · Physics 2025-11-11 Jan Rozman , Sumesh P. Thampi , Julia M. Yeomans

We present a hydrodynamic theory to describe shear flows in developing epithelial tissues. We introduce hydrodynamic fields corresponding to state properties of constituent cells as well as a contribution to overall tissue shear flow due to…

Active materials are capable of converting free energy into directional motion, giving rise to striking dynamical phenomena. Developing a general understanding of their structure in relation to the underlying non-equilibrium physics would…

Soft Condensed Matter · Physics 2018-10-17 Nitin Kumar , Rui Zhang , Juan J. de Pablo , Margaret L. Gardel

We analyze the phase behavior of lyotropic nematic liquid crystals in the self-organizing flow, viz. so called active nematics (AN). Their elastic properties are mutually caused by evolution of topological defects (for instance,…

Soft Condensed Matter · Physics 2018-02-20 L. V. Elnikova

The study of liquid crystals at equilibrium has led to fundamental insights into the nature of ordered materials, as well as to practical applications such as display technologies. Active nematics are a fundamentally different class of…

Soft Condensed Matter · Physics 2015-08-19 Stephen J. DeCamp , Gabriel S. Redner , Aparna Baskaran , Michael F. Hagan , Zvonimir Dogic

Monolayers of anisotropic cells exhibit long-ranged orientational order and topological defects. During the development of organisms, orientational order often influences morphogenetic events. However, the linkage between the mechanics of…

Soft Condensed Matter · Physics 2021-01-20 Carles Blanch-Mercader , Pau Guillamat , Aurélien Roux , Karsten Kruse

Topological defects in systems with liquid-crystalline order are crucial in determining their large-scale properties. In active systems, they are known to have properties impossible at equilibrium: for example, $+1/2$ defects in…

Soft Condensed Matter · Physics 2026-02-17 Giacomo Marco La Montagna , Sumeja Burekovic , Ananyo Maitra , Cesare Nardini

We formulate the statistical dynamics of topological defects in the active nematic phase, formed in two dimensions by a collection of self-driven particles on a substrate. An important consequence of the non-equilibrium drive is the…

Soft Condensed Matter · Physics 2018-09-12 Suraj Shankar , Sriram Ramaswamy , M. Cristina Marchetti , Mark J. Bowick

We investigate the mechanics of an active droplet endowed with internal nematic order and surrounded by an isotropic Newtonian fluid. Using numerical simulations we demonstrate that, due to the interplay between the active stresses and the…

Soft Condensed Matter · Physics 2014-04-23 L. Giomi , A. DeSimone

Active matter encompasses different nonequilibrium systems in which individual constituents convert energy into non-conservative forces or motion at the microscale. This review provides an elementary introduction to the role of topology in…

Soft Condensed Matter · Physics 2022-06-28 Suraj Shankar , Anton Souslov , Mark J. Bowick , M. Cristina Marchetti , Vincenzo Vitelli

Biological cells can actively tune their intracellular architecture according to their overall shape. Here we explore the rheological implication of such coupling in a minimal model of a dense cellular material where each cell exerts an…

Soft Condensed Matter · Physics 2022-04-13 Shao-Zhen Lin , Matthias Merkel , Jean-François Rupprecht

We study the dynamics of topological defects in active nematic films with spatially-varying activity and consider two setups: i) a constant activity gradient, and ii) a sharp jump in activity. A constant gradient of extensile (contractile)…

Soft Condensed Matter · Physics 2022-09-02 Jonas Rønning , M. Cristina Marchetti , Luiza Angheluta

We study two models of overdamped self-propelled disks in two dimensions, with and without aligning interactions. Active mesoscale flows leading to chaotic advection emerge in both models in the homogeneous dense fluid away from dynamical…

Soft Condensed Matter · Physics 2024-06-07 Yann-Edwin Keta , Juliane Klamser , Robert L. Jack , Ludovic Berthier

Topological defects play a central role in the physics of many materials, including magnets, superconductors and liquid crystals. In active fluids, defects become autonomous particles that spontaneously propel from internal active stresses…

Soft Condensed Matter · Physics 2024-06-12 Suraj Shankar , Luca V. D. Scharrer , Mark J. Bowick , M. Cristina Marchetti

Collective cell migration governs a range of physiological and pathological processes, from tissue morphogenesis to cancer invasion, in which topological defects arise as an inevitable consequence of frequent cellular rearrangement and…

Soft Condensed Matter · Physics 2026-04-01 Jiusi Zhang , Chung Wing Chan , Bo Li , Rui Zhang

Active matter is naturally out of equilibrium which results in the emergence of diverse dynamic steady states, including the omnipresent chaotic state known as the active turbulence. However, much less is known how active systems…

Soft Condensed Matter · Physics 2023-04-07 Nika Kralj , Miha Ravnik , Žiga Kos

We introduce a general description of localised distortions in active nematics using the framework of active nematic multipoles. We give the Stokesian flows for arbitrary multipoles in terms of differentiation of a fundamental flow response…

Soft Condensed Matter · Physics 2023-04-04 Alexander J. H. Houston , Gareth P. Alexander

Active matter is characterized by its ability to induce motion by self-generated stress. In the case of a solid, such motion can lead to shape transformations. The stress-generating components can be anisotropic endowing the material with…

Soft Condensed Matter · Physics 2022-05-16 D. J. G. Pearce , S. Gat , G. Livne , A. Bernheim-Groswasser , K. Kruse