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Many biological systems consist of self-motile and passive agents both of which contribute to overall functionality. However, there are very few studies of the properties of such mixtures. Here we formulate a model for mixtures of…

Soft Condensed Matter · Physics 2015-05-25 Denis F. Hinz , Alexander Panchenko , Tae-Yeon Kim , Eliot Fried

The motility-induced phase separation (MIPS) phenomenon in active matter has been of great interest for the past decade or so. A central conceptual puzzle is that this behavior, which is generally characterized as a nonequilibrium…

Statistical Mechanics · Physics 2024-05-28 Venkat Venkatasubramanian , Abhishek Sivaram , N. Sanjeevrajan , Arun Sankar

The field of active matter explores the behaviors of self propelled agents out of equilibrium, with active suspensions, such as swimming bacteria in solutions, serving as impactful models. These systems exhibit spatio-temporal patterns akin…

Soft Condensed Matter · Physics 2025-08-26 Pratikshya Jena , Shradha Mishra

Motility and nonreciprocity are two primary mechanisms for self-organization in active matter. In a recent study [Phys. Rev. Lett. 131, 148301 (2023)], we explored their joint influence in a minimal model of two-species quorum-sensing…

Soft Condensed Matter · Physics 2025-04-15 Yu Duan , Jaime Agudo-Canalejo , Ramin Golestanian , Benoît Mahault

We study non-equilibrium phases for interacting two-dimensional self-propelled particles with isotropic pair-wise interactions using a persistent kinetic Monte Carlo (MC) approach. We establish the quantitative phase diagram, including the…

Soft Condensed Matter · Physics 2019-04-17 Juliane U. Klamser , Sebastian C. Kapfer , Werner Krauth

We consider a mixture of passive (i.e., Brownian) and active (e.g., bacterial or colloidal swimmers) particles, and analyze the stability conditions of either uniformly mixed or phase segregated steady states consisting of phases enriched…

Statistical Mechanics · Physics 2015-09-21 Alexander Y. Grosberg , Jean-François Joanny

Mixtures of microtubules and molecular motors form active materials with diverse dynamical behaviors that vary based on their constituents' molecular properties. We map the non-equilibrium phase diagram of microtubules and tip-accumulating…

Soft Condensed Matter · Physics 2021-08-04 Bezia Lemma , Noah P. Mitchell , Radhika Subramanian , Daniel J. Needleman , Zvonimir Dogic

Motivated by recent experiments, we study a system of self-propelled colloids that experience short-range attractive interactions and are confined to a surface. Using simulations we find that the phase behavior for such a system is…

Soft Condensed Matter · Physics 2013-07-30 Gabriel S. Redner , Aparna Baskaran , Michael F. Hagan

Motility-induced phase separation (MIPS) is a well-studied nonequilibrium collective phenomenon observed in active particles. Recently, there has been growing interest in how coupling the self-propulsion of active particles to chemical…

Soft Condensed Matter · Physics 2025-07-14 Euijoon Kwon , Yongjae Oh , Yongjoo Baek

Biomolecular condensates in cells are often rich in catalytically-active enzymes. This is particularly true in the case of the large enzymatic complexes known as metabolons, which contain different enzymes that participate in the same…

Subcellular Processes · Quantitative Biology 2021-10-20 Vincent Ouazan-Reboul , Jaime Agudo-Canalejo , Ramin Golestanian

Materials undergoing both phase separation and chemical reactions (defined here as all processes that change particle type or number) form an important class of non-equilibrium systems. Examples range from suspensions of self-propelled…

Soft Condensed Matter · Physics 2020-09-21 Yuting I. Li , Michael E. Cates

We computationally study suspensions of slow and fast active Brownian particles that have undergone motility induced phase separation and are at steady state. Such mixtures, of varying non-zero activity, remain largely unexplored even…

Soft Condensed Matter · Physics 2024-07-11 Nicholas J Lauersdorf , Ehssan Nazockdast , Daphne Klotsa

Turing's mechanism is often invoked to explain periodic patterns in nature, although direct experimental support is scarce. Turing patterns form in reaction-diffusion systems when the activating species diffuse much slower than the…

Biological Physics · Physics 2024-03-15 Lucas Menou , Chengjie Luo , David Zwicker

Meso-scale turbulence was originally observed experimentally in various suspensions of swimming bacteria, as well as in the collective motion of active colloids. The corresponding large-scale dynamical patterns were reproduced in a simple…

Soft Condensed Matter · Physics 2021-03-30 Vasco M. Worlitzer , Gil Ariel , Avraham Be'er , Holger Stark , Markus Bär , Sebastian Heidenreich

Non-equilibrium phase separating systems with reactions can break time-reversal symmetry (TRS) in two distinct ways. Firstly, the conservative and non-conservative sectors of the dynamics can be governed by incompatible free energies; when…

Soft Condensed Matter · Physics 2021-05-31 Yuting I. Li , Michael E. Cates

Dry active matter systems are well-known to exhibit Motility-Induced Phase Separation (MIPS). However, in wet active systems, attractive hydrodynamic interactions mediated by active particles stalled at a boundary can introduce…

Soft Condensed Matter · Physics 2026-01-01 Kingshuk Panja , Rajesh Singh

Active colloids exhibit persistent motion, which can lead to motility-induced phase separation (MIPS). However, there currently exists no microscopic theory to account for this phenomenon. We report a first-principles theory, free of fit…

Soft Condensed Matter · Physics 2015-06-24 T. F. F. Farage , P. Krinninger , J. M. Brader

The transport properties of colloidal particles in active liquids have been studied extensively. It has led to a deeper understanding of the interactions between passive and active particles. However, the phase behavior of colloidal…

Soft Condensed Matter · Physics 2023-01-30 Pragya Kushwaha , Vivek Semwal , Sayan Maity , Shraddha Mishra , Vijayakumar Chikkadi

The intestinal microbiota plays important roles in digestion and resistance against entero-pathogens. As with other ecosystems, its species composition is resilient against small disturbances but strong perturbations such as antibiotics can…

Quantitative Methods · Quantitative Biology 2015-06-04 Vanni Bucci , Serena Bradde , Giulio Biroli , Joao B. Xavier

Collectives of actively-moving particles can spontaneously separate into dilute and dense phases -- a fascinating phenomenon known as motility-induced phase separation (MIPS). MIPS is well-studied for randomly-moving particles with no…

Biological Physics · Physics 2023-01-31 Hongbo Zhao , Andrej Košmrlj , Sujit S. Datta