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Explaining how competing species coexist remains a central question in ecology. The well-known competitive exclusion principle (CEP) states that two species competing for the same resource cannot stably coexist, and more generally, that the…

Biological Physics · Physics 2026-01-01 Wei Tao , Ju Kang , Wenxiu Yang , Yiyuan Niu , Xin Wang

Explaining biodiversity is a central focus in theoretical ecology. A significant obstacle arises from the Competitive Exclusion Principle (CEP), which states that two species competing for the same type of resources cannot coexist at…

Populations and Evolution · Quantitative Biology 2025-06-18 Ju Kang , Yiyuan Niu , Xin Wang

Explaining biodiversity in nature is a fundamental problem in ecology. An outstanding challenge is embodied in the so-called Competitive Exclusion Principle: two species competing for one limiting resource cannot coexist at constant…

Populations and Evolution · Quantitative Biology 2020-04-13 Xin Wang , Yang-Yu Liu

In apparent contradiction to competition theory, the number of known, co-existing plankton species far exceeds their explicable biodiversity - a discrepancy termed the Paradox of the Plankton. We introduce a new game-theoretic model for…

Populations and Evolution · Quantitative Biology 2020-12-14 Susanne Menden-Deuer , Julie Rowlett

Understanding the mechanisms that sustain high biodiversity remains a central challenge. MacArthur's classical consumer-resource model (MCRM) suggests that consumer diversity is limited by the number of available resources, yet empirical…

Populations and Evolution · Quantitative Biology 2025-08-25 Seong-Gyu Yang , Hye Jin Park

What determines biodiversity in nature is a prominent issue in ecology, especially in biotic resource systems that are typically devoid of cross-feeding. Here, we show that by incorporating pairwise encounters among consumer individuals…

Populations and Evolution · Quantitative Biology 2025-06-18 Ju Kang , Shijie Zhang , Yiyuan Niu , Xin Wang

Local coexistence of species in large ecosystems is traditionally explained within the broad framework of niche theory. However, its rationale hardly justifies rich biodiversity observed in nearly homogeneous environments. Here we consider…

Populations and Evolution · Quantitative Biology 2021-11-17 Deepak Gupta , Stefano Garlaschi , Samir Suweis , Sandro Azaele , Amos Maritan

In its simplest form, the competitive exclusion principle states that a number of species competing for a smaller number of resources cannot coexist. However, it has been observed empirically that in some settings it is possible to have…

Populations and Evolution · Quantitative Biology 2021-02-18 Alexandru Hening , Dang H. Nguyen

Species introductions to new habitats can cause a decline in the population size of competing native species and consequently also in their genetic diversity. We are interested in why these adverse effects are weak in some cases whereas in…

Populations and Evolution · Quantitative Biology 2013-01-03 Meike J. Wittmann , Martin Hutzenthaler , Wilfried Gabriel , Dirk Metzler

This is the first of two papers where we discuss the limits imposed by competition to the biodiversity of species communities. In this first paper we study the coexistence of competing species at the fixed point of population dynamic…

Populations and Evolution · Quantitative Biology 2007-05-23 Ugo Bastolla , Michael Lässig , Susanna C. Manrubia , Angelo Valleriani

The processes and mechanisms underlying the origin and maintenance of biological diversity have long been of central importance in ecology and evolution. The competitive exclusion principle states that the number of coexisting species is…

Populations and Evolution · Quantitative Biology 2020-12-02 Michael Doebeli , Eduardo Cancino Jaque , Iaroslav Ispolatov

This is the second of two papers dedicated to the relationship between population models of competition and biodiversity. Here we consider species assembly models where the population dynamics is kept far from fixed points through the…

Populations and Evolution · Quantitative Biology 2007-05-23 Ugo Bastolla , Michael Lässig , Susanna C. Manrubia , Angelo Valleriani

How large ecosystems can create and maintain the remarkable biodiversity we see in nature is probably one of the biggest open questions in science, attracting attention from different fields, from Theoretical Ecology to Mathematics and…

Populations and Evolution · Quantitative Biology 2023-01-03 Violeta Calleja-Solanas , Nagi Khalil , Jesús Gómez-Gardeñes , Emilio Hernández-García , Sandro Meloni

Ecosystems are commonly organized into trophic levels -- organisms that occupy the same level in a food chain (e.g., plants, herbivores, carnivores). A fundamental question in theoretical ecology is how the interplay between trophic…

Populations and Evolution · Quantitative Biology 2023-03-07 Zhijie Feng , Robert Marsland , Jason W. Rocks , Pankaj Mehta

Explaining coexistence in species-rich communities of primary producers remains a challenge for ecologists because of their likely competition for shared resources. Following Hutchinson's seminal suggestion, many theoreticians have tried to…

Populations and Evolution · Quantitative Biology 2019-04-25 Coralie Picoche , Frederic Barraquand

The competitive exclusion principle asserts that coexisting species must occupy distinct ecological niches (i.e. the number of surviving species can not exceed the number of resources). An open question is to understand if and how different…

Populations and Evolution · Quantitative Biology 2020-07-29 Wenping Cui , Robert Marsland , Pankaj Mehta

Resource competition is a fundamental interaction in natural communities.However little is known about competition in spatial environments where organisms are able to regulate resource distributions. Here, we analyze the competition of two…

Populations and Evolution · Quantitative Biology 2011-02-24 Alexei B. Ryabov , Bernd Blasius

Ecosystems frequently display the coexistence of diverse species under resource competition, typically resulting in skewed distributions of rarity and abundance. A potential driver of such coexistence is environmental fluctuations that…

Populations and Evolution · Quantitative Biology 2025-08-07 Davide Zanchetta , Deepak Gupta , Sofia Moschin , Samir Suweis , Amos Maritan , Sandro Azaele

Given an endogenous timescale set by invasion in a constant environment, we introduced periodic temporal variation in competitive superiority by alternating the species' propagation rates. By manipulating habitat size and introduction rate,…

Populations and Evolution · Quantitative Biology 2011-05-10 Lauren O'Malley , G. Korniss , Sai Satya Praveen Mungara , Thomas Caraco

Biodiversity widely observed in ecological systems is attributed to the dynamical balance among the competing species. The time-varying populations of the interacting species are often captured rather well by a set of deterministic…

Populations and Evolution · Quantitative Biology 2010-05-25 Yen-Chih Lin , Tzay-Ming Hong , Hsiu-Hau Lin
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