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Microswimmers (planktonic microorganisms or artificial active particles) immersed in a fluid interact with the ambient flow, altering their trajectories. In surface gravity waves, a common goal for microswimmers is vertical migration (e.g.,…

Soft Condensed Matter · Physics 2022-03-01 Kunlin Ma , Nimish Pujara , Jean-Luc Thiffeault

Squirmers are models of a class of microswimmers, such as ciliated organisms and phoretic particles, that self-propel in fluids without significant deformation of their body shape. Available techniques for their simulation are based on the…

Numerical Analysis · Mathematics 2019-03-20 Stevens Paz Sánchez , Gustavo C. Buscaglia

Flagellated microswimmers are ubiquitous in natural habitats. Understanding the hydrodynamic behavior of these cells is of paramount interest, owing to their applications in bio-medical engineering and disease spreading. Since the last two…

Fluid Dynamics · Physics 2024-05-20 H. Gidituri , G. Kabacaoğlu , M. Ellero , F. Balboa Usabiaga

We analyse a simple 'Stokesian squirmer' model for the enhanced mixing due to swimming micro-organisms. The model is based on a calculation of Thiffeault & Childress [Physics Letters A, 374, 3487 (2010), arXiv:0911.5511], where fluid…

Fluid Dynamics · Physics 2013-09-24 Zhi Lin , Jean-Luc Thiffeault , Stephen Childress

We study two microswimmers consisting of a spherical rigid head and a passive elastic tail. In the first one the tail is clamped to the head, and the system oscillates under the action of an external torque. In the second one, head and tail…

Fluid Dynamics · Physics 2017-02-02 Giancarlo Cicconofri , Antonio DeSimone

Self-propelled particles have been experimentally shown to orbit spherical obstacles and move along surfaces. Here, we theoretically and numerically investigate this behavior for a hydrodynamic squirmer interacting with spherical objects…

Soft Condensed Matter · Physics 2019-07-05 Michael Kuron , Philipp Stärk , Christian Holm , Joost de Graaf

Biological and artificial microswimmers often have to propel through a variety of environments, ranging from heterogeneous suspending media to strong geometrical confinement. Under confinement, local flow fields generated by microswimmers,…

Soft Condensed Matter · Physics 2024-06-04 Florian A. Overberg , Gerhard Gompper , Dmitry A. Fedosov

We combine a general formulation of microswimmmer equations of motion with a numerical bead-shell model to calculate the hydrodynamic interactions with the fluid, from which the swimming speed, power and efficiency are extracted. From this…

Soft Condensed Matter · Physics 2017-03-07 Bram Bet , Gijs Boosten , Marjolein Dijkstra , René van Roij

The spatiotemporal dynamics in systems of active self-propelled particles is controlled by the propulsion mechanism in combination with various direct interactions, such as steric repulsion, hydrodynamics, and chemical fields. Yet, these…

Soft Condensed Matter · Physics 2024-06-03 Mario Theers , Elmar Westphal , Kai Qi , Roland G. Winkler , Gerhard Gompper

Biological microswimmers such as bacteria and sperm cells often encounter complex biological fluid environments. Here we use the well-known squirmer microswimmer model to show the importance of the local fluid microstructure and…

Soft Condensed Matter · Physics 2023-07-19 Andreas Zöttl

Biological microswimmers alter their motility in complex corner geometries, facilitating their survival. However, the dynamical features of low-Reynolds-number swimming at corners remain undefined. Here, we use active droplet microswimmers…

Soft Condensed Matter · Physics 2026-04-22 Subhasish Guchhait , Harshita Tiwari , Sumesh P. Thampi , Ranabir Dey

We study the dynamics of gyrotactic microswimmers suspended in homogeneous and isotropic turbulence by using direct numerical simulations (DNS). The swimmers are characterized by three non-dimensional parameters: their aspect ratio…

Soft Condensed Matter · Physics 2026-04-24 Suraj Kumar Nayak , Vishwanath Shukla , Akshay Bhatnagar

Pairwise hydrodynamic interactions of microswimmers form the fundamental building blocks for understanding their more complex collective behaviors. In this work, we revisit the canonical problem of two interacting squirmers swimming along…

Inspired by the classical Kepler and Rutherford problem, we investigate an analogous set-up in the context of active microswimmers: the behavior of a deformable microswimmer in a swirl flow. First we identify new steady bound states in the…

Soft Condensed Matter · Physics 2014-09-11 Mitsusuke Tarama , Andreas M. Menzel , Hartmut Löwen

Microorganisms ofter move in confined, disordered environments, where hydrodynamic couplings can modify their transport behavior. Using extensive finite-element simulations, we investigate the dynamics of microswimmers -- modeled as…

Soft Condensed Matter · Physics 2026-03-24 Mirko Residori , Sebastian Aland , Christina Kurzthaler

Bacteria in biofilms form complex structures and can collectively migrate within mobile aggregates, which is referred to as swarming. This behavior is influenced by a combination of various factors, including morphological characteristics…

Soft Condensed Matter · Physics 2024-01-19 Bohan Wu-Zhang , Dmitry A. Fedosov , Gerhard Gompper

Microswimmers in suspension exhibit collective swimming behaviour, forming various self-organised structures including ordered, aggregated, and turbulent-like structures. When mixed with passive particles phase-separation is known to occur,…

Fluid Dynamics · Physics 2026-03-26 Alexander Chamolly , Takuji Ishikawa

Unlike macroscopic swimmers, microswimmers operate in a low-Reynolds-number regime dominated by viscous forces. This paper investigates the controllability of a magnetic microswimmer composed of a spherical magnetic head and an elastic,…

Optimization and Control · Mathematics 2025-11-05 Lucas Palazzolo , Mickaël Binois , Laëtitia Giraldi

We present a lattice Boltzmann study of the hydrodynamics of a fully resolved squirmer, radius R, confined in a slab of fluid between two no-slip walls. We show that the coupling between hydrodynamics and short-range repulsive interactions…

Soft Condensed Matter · Physics 2016-03-30 Juho S. Lintuvuori , Aidan T. Brown , Kevin Stratford , Davide Marenduzzo

We find the mixing induced by a small swimming organism in a density stratified fluid. We model the swimmer as a spherical squirmer and quantify mixing through a mixing efficiency that is the ratio of rate of change of potential energy of…

Soft Condensed Matter · Physics 2025-02-26 Vaseem A. Shaik , Gwynn J. Elfring