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Population heterogeneity is ubiquitous among active living systems, but little is known about its role in determining their spatial organization and large-scale dynamics. Combining evidence from synthetic active fluids assembled from…

Soft Condensed Matter · Physics 2023-11-27 Samadarshi Maity , Alexandre Morin

We propose a hydrodynamic theory to examine the emergence of contraction waves in dense active liquids composed of pulsating deformable particles. Our theory couples the liquid density with a chemical phase that determines the periodic…

Statistical Mechanics · Physics 2025-09-24 Tirthankar Banerjee , Thibault Desaleux , Jonas Ranft , Étienne Fodor

The microalga Chlamydomonas Reinhardtii (CR) is used here as a model system to study the effect of complex environments on the swimming of micro-organisms. Its motion can be modelled by a run and tumble mechanism so that it describes a…

Soft Condensed Matter · Physics 2019-03-27 Marvin Brun-Cosme-Bruny , Eric Bertin , Benoît Coasne , Philippe Peyla , Salima Rafaï

Organisms often swim through density stratified fluids. In this Letter, we investigate the dynamics of small active particles swimming in density gradients and report theoretical evidence of taxis as a result of density stratification…

Soft Condensed Matter · Physics 2023-08-02 Vaseem A. Shaik , Gwynn J. Elfring

There is increasing evidence that mammalian cells not only crawl on substrates but can also swim in fluids. To elucidate the mechanisms of the onset of motility of cells in suspension, a model which couples actin and myosin kinetics to…

Biological Physics · Physics 2019-09-18 Alexander Farutin , Jocelyn Etienne , Chaouqi Misbah , Pierre Recho

Recently, there is much interest in droplet condensation on soft or liquid/liquid-like substrates. Droplets can deform soft and liquid interfaces resulting in a wealth of phenomena not observed on hard, solid surfaces (e.g., increased…

Soft Condensed Matter · Physics 2023-11-14 Marcus Lin , Philseok Kim , Sankara Arunachalam , Rifan Hardian , Solomon Adera , Joanna Aizenberg , Xi Yao , Dan Daniel

We analyse collective motion that occurs during rare (large deviation) events in systems of active particles, both numerically and analytically. We discuss the associated dynamical phase transition to collective motion, which occurs when…

Statistical Mechanics · Physics 2021-02-10 Yann-Edwin Keta , Étienne Fodor , Frédéric van Wijland , Michael E. Cates , Robert L. Jack

We derive from first principles a three-dimensional theory of self-propelled particle swarming in a viscous fluid environment. Our model predicts emergent collective behavior that depends critically on fluid opacity, mechanism of…

Soft Condensed Matter · Physics 2026-05-12 Yao-Li Chuang , M. R. D'Orsogna , T. Chou

We propose a model for the motion of a single active particle in a heterogeneous environment where the heterogeneity may arise due to the crowding, conformational fluctuations and/or slow rearrangement of the surroundings. Describing the…

Statistical Mechanics · Physics 2022-03-04 Koushik Goswami , Rajarshi Chakrabarti

We study the activity of "living" droplets, which confine 1-6 mesoswimmers in 3D using a superhydrophobic substrate. The swimmers induce oscillations of the droplets at their inherent resonant frequencies, regardless of swimmer size and…

Soft Condensed Matter · Physics 2026-01-30 L. Malik , N. Sharadhi , M. Lamminmäki , R. A. Lara , V. Jokinen , M. Backholm

Active systems comprising micron-sized self-propelling units, also termed microswimmers, are promising candidates for the bottom-up assembly of small structures and reconfigurable materials. Here we leverage field-driven colloidal assembly…

Soft Condensed Matter · Physics 2024-12-24 Carolina van Baalen , Laura Alvarez , Robert Style , Lucio Isa

Motility is a fundamental survival strategy of bacteria to navigate porous environments. Swimming cells thrive in quiescent wetlands and sediments at the bottom of the marine water column, where they mediate many essential biogeochemical…

Soft Condensed Matter · Physics 2022-01-11 Amin Dehkharghani , Nicolas Waisbord , Jeffrey S. Guasto

We develop a minimal hydrodynamic model, without an orientational order parameter, for assemblies of contractile swimmers encapsulated in a droplet of a binary-fluid emulsion. Our model uses two coupled scalar order parameters, $\phi$ and…

Soft Condensed Matter · Physics 2022-09-13 Nadia Bihari Padhan , Rahul Pandit

Active (i.e., self-propelled or swimming) particles moving through an isotropic fluid exhibit conventional diffusive behavior. We report anomalous diffusion of an active particle moving in an anisotropic, nematic background. Whilst the…

Soft Condensed Matter · Physics 2016-06-22 John Toner , Hartmut Löwen , Henricus H. Wensink

Systems of motile microorganisms exhibit a multitude of collective phenomena, including motility-induced phase separation and turbulence. Sensing of the environment and adaptation of movement plays an essential role in the emergent…

Soft Condensed Matter · Physics 2024-06-26 Segun Goh , Elmar Westphal , Roland G. Winkler , Gerhard Gompper

Active systems, from bacterial suspensions to cellular monolayers, are continuously driven out of equilibrium by local injection of energy from their constituent elements and exhibit turbulent-like and chaotic patterns. Here we demonstrate…

Soft Condensed Matter · Physics 2016-02-04 Amin Doostmohammadi , Michael F. Adamer , Sumesh P. Thampi , Julia M. Yeomans

Living systems are made of active materials with microscopic components that work together to perform macroscopic biological tasks. The breakdown of these collective functionalities leads to diseases, which, conversely, could be treated by…

Active matter comprises individual units that convert energy into mechanical motion. In many examples, such as bacterial systems and biofilament assays, constituent units are elongated and can give rise to local nematic orientational order.…

Soft Condensed Matter · Physics 2020-11-17 He Li , Xia-qing Shi , Mingji Huang , Xiao Chen , Mingfeng Xiao , Chenli Liu , Hugues Chate , H. P. Zhang

In systems as diverse as migrating mammals to road traffic, crowding acts to inhibit efficient collective movement. Bacteria, however, are observed to move in very dense groups containing billions of individuals without causing the gridlock…

We show that "dry" active nematics, e.g. collections of shaken elongated granular particles, exhibit large-scale spatiotemporal chaos made of interacting dense, ordered, band-like structures in a parameter region including the linear onset…

Soft Condensed Matter · Physics 2014-09-16 Sandrine Ngo , Anton Peshkov , Igor S. Aranson , Eric Bertin , Francesco Ginelli , Hugues Chaté