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Three-dimensional simulations with fully resolved hydrodynamics are performed to study the collective motion of model swimmers in confinement. We show that certain swimming mechanisms can lead to traveling wave-like collective motion even…

Soft Condensed Matter · Physics 2016-04-20 Norihiro Oyama , John Jairo Molina , Ryoichi Yamamoto

Many microswimmers are inherently chiral, and this chirality can introduce fascinating behaviors in a collection of microswimmers. The dynamics become even more intriguing when two types of microswimmers with distinct chirality are mixed.…

Soft Condensed Matter · Physics 2025-09-03 Divya Kushwaha , Shradha Mishra

Cells swimming in confined environments are attracted by surfaces. We measure the steady-state distribution of smooth-swimming bacteria (Escherichia coli) between two glass plates. In agreement with earlier studies, we find a strong…

Soft Condensed Matter · Physics 2008-10-02 Allison P. Berke , Linda Turner , Howard C. Berg , Eric Lauga

Active matter systems encompass both natural and artificially created systems consisting of numerous active particles. These particles actively consume energy to propel themselves or exert mechanical forces, leading to intricate behaviors…

Statistical Mechanics · Physics 2026-05-12 Mintu Karmakar

Active bodies in viscous fluids interact hydrodynamically through self-generated flows. Here we study spontaneous aggregation induced by hydrodynamic flow in a suspension of stiff, apolar, active filaments. Lateral hydrodynamic attractions…

Soft Condensed Matter · Physics 2015-08-20 Ankita Pandey , P. B. Sunil Kumar , R. Adhikari

Active flexible filaments form the classical continuum framework for modelling the locomotion of spermatozoa and algae driven by the periodic oscillation of flagella. This framework also applies to the locomotion of various artificial…

Biological Physics · Physics 2025-10-14 John Severn , Eric Lauga

Capturing the emergence of deformation waves in contractile living tissues is a challenge that has recently been tackled with models of actively deformable particles. Inspired by the anisotropic deformation of cardiomyocytes in cardiac…

Soft Condensed Matter · Physics 2026-05-26 Luca Casagrande , Alessandro Manacorda , Etienne Fodor

When interacting motile units self-organize into flocks, they realize one of the most robust ordered state found in nature. However, after twenty five years of intense research, the very mechanism controlling the ordering dynamics of both…

Soft Condensed Matter · Physics 2021-10-04 Amélie Chardac , Ludwig A. Hoffmann , Yoann Poupart , Luca Giomi , Denis Bartolo

Micron-sized self-propelled (active) particles can be considered as model systems for characterizing more complex biological organisms like swimming bacteria or motile cells. We produce asymmetric microswimmers by soft lithography and study…

Here we show that coupling to curvature has profound effects on collective motion in active systems, leading to patterns not observed in flat space. Biological examples of such active motion in curved environments are numerous: curvature…

Soft Condensed Matter · Physics 2015-03-05 Rastko Sknepnek , Silke Henkes

Near a solid boundary, E. coli swims in clockwise circular motion. We provide a hydrodynamic model for this behavior. We show that circular trajectories are natural consequences of force-free and torque-free swimming, and the hydrodynamic…

Soft Condensed Matter · Physics 2016-08-31 Eric Lauga , Willow R. DiLuzio , George M. Whitesides , Howard A. Stone

Active matter is not only indispensable to our understanding of diverse biological processes, but also provides a fertile ground for discovering novel physics. Many emergent properties impossible for equilibrium systems have been…

Soft Condensed Matter · Physics 2022-05-27 Thibault Bertrand , Chiu Fan Lee

Active fluids exhibit spontaneous flows with complex spatiotemporal structure, which have been observed in bacterial suspensions, sperm cells, cytoskeletal suspensions, self-propelled colloids, and cell tissues. Despite occurring in the…

Soft Condensed Matter · Physics 2022-10-05 Ricard Alert , Jaume Casademunt , Jean-François Joanny

Self-propulsion of cellular microswimmers generates flow signatures, commonly classified as pusher- and puller-type, which characterize hydrodynamic interactions with other cells or boundaries. Using experimentally measured beat patterns,…

Cell Behavior · Quantitative Biology 2016-01-20 Gary S. Klindt , Benjamin M. Friedrich

Active diffusiophoresis - swimming through interaction with a self-generated, neutral, solute gradient - is a paradigm for autonomous motion at the micrometer scale. We study this propulsion mechanism within a linear response theory.…

Statistical Mechanics · Physics 2015-05-30 Benedikt Sabass , Udo Seifert

Swimming microorganisms can influence the diffusion of passive particles. The effect of this swimmer-particle interaction depends on different properties, such as the hydrodynamic field of the swimmer and the relative sizes of…

Soft Condensed Matter · Physics 2020-05-08 Florian von Rüling , Francine Kolley , Alexey Eremin

We investigate density fluctuations in three-dimensional chiral active fluids by using a simple model of helical self-propelled particles. Helical motion is generated by a constant angular velocity (or chiral torque) acting on the…

Soft Condensed Matter · Physics 2025-10-29 Yuta Kuroda , Takeshi Kawasaki , Kunimasa Miyazaki

Purcell's scallop theorem states that swimmers deforming their shapes in a time-reversible manner ("reciprocal" motion) cannot swim. Using numerical simulations and theoretical calculations we show here that in a fluctuating environment,…

Soft Condensed Matter · Physics 2011-08-30 Eric Lauga

The spherical alga Volvox swims by means of flagella on thousands of surface somatic cells. This geometry and its large size make it a model organism for studying the fluid dynamics of multicellularity. Remarkably, when two nearby Volvox…

A suspension of gyrotactic microalgae Chlamydomonas augustae swimming in a cylindrical water vessel in solid-body rotation is studied. Our experiments show that swimming algae form an aggregate around the axis of rotation, whose intensity…

Fluid Dynamics · Physics 2015-11-05 M. Cencini , M. Franchino , F. Santamaria , G. Boffetta