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The survival of many microorganisms, like \textit{Leptospira} or \textit{Spiroplasma} bacteria, can depend on their ability to navigate towards regions of favorable viscosity. While this ability, called viscotaxis, has been observed in…

Soft Condensed Matter · Physics 2018-05-23 Benno Liebchen , Paul Monderkamp , Borge ten Hagen , Hartmut Löwen

Biological microswimmers alter their motility in complex corner geometries, facilitating their survival. However, the dynamical features of low-Reynolds-number swimming at corners remain undefined. Here, we use active droplet microswimmers…

Soft Condensed Matter · Physics 2026-04-22 Subhasish Guchhait , Harshita Tiwari , Sumesh P. Thampi , Ranabir Dey

Inspired by the classical Kepler and Rutherford problem, we investigate an analogous set-up in the context of active microswimmers: the behavior of a deformable microswimmer in a swirl flow. First we identify new steady bound states in the…

Soft Condensed Matter · Physics 2014-09-11 Mitsusuke Tarama , Andreas M. Menzel , Hartmut Löwen

Swimming cells and microorganisms must often move though complex fluids that contain an immersed microstructure such as polymer molecules, or filaments. In many important biological processes, such as mammalian reproduction and bacterial…

Fluid Dynamics · Physics 2018-08-06 Arshad Kamal , Eric E Keaveny

We suggest several reciprocal swimming mechanisms that lead to a locomotion only in viscoelastic fluids. The first situation is to have a difference between the two amplitudes of the oscillatory arm motion for a three-sphere microswimmer.…

Soft Condensed Matter · Physics 2021-06-08 Kento Yasuda , Mizuki Kuroda , Shigeyuki Komura

In the study of microscopic flows, self-propulsion has been particularly topical in recent years, with the rise of miniature artificial swimmers as a new tool for flow control, low Reynolds number mixing, micromanipulation or even drug…

Geometric confinements are frequently encountered in soft matter systems and in particular significantly alter the dynamics of swimming microorganisms in viscous media. Surface-related effects on the motility of microswimmers can lead to…

The behaviour of microscopic swimmers has previously been explored near large scale confining geometries and in the presence of very small-scale surface roughness. Here we consider an intermediate case of how a simple microswimmer, the…

Fluid Dynamics · Physics 2021-09-14 Kenta Ishimoto , Eamonn A. Gaffney , David J. Smith

From bacteria and sperm cells to artificial microrobots, self-propelled microscopic objects at low Reynolds numbers often perceive fluctuating mechanical and chemical stimuli and contact exterior wall boundaries both in nature and the…

Soft Condensed Matter · Physics 2023-12-22 Yoshiki Hiruta , Kenta Ishimoto

Biological and artificial microswimmers often self-propel in external flows of vortical nature; relevant examples include algae in small-scale ocean eddies, spermatozoa in uterine peristaltic flows and bacteria in microfluidic devices. A…

Biological Physics · Physics 2022-11-14 Ivan Tanasijevic , Eric Lauga

We perform hydrodynamic simulations using the method of multi-particle collision dynamics and a theoretical analysis to study a single squirmer microswimmer at high P\'eclet number, which moves in a low Reynolds number fluid and under…

Fluid Dynamics · Physics 2018-03-14 Felix Rühle , Johannes Blaschke , Jan-Timm Kuhr , Holger Stark

Microswimmers typically move near walls, which can strongly influence their motion. However, direct experimental measurements of swimmer-wall separation remain elusive to date. Here, we determine this separation for model catalytic…

Soft Condensed Matter · Physics 2020-12-30 Stefania Ketzetzi , Joost de Graaf , Daniela J. Kraft

Microswimmers are often found in heterogeneous and crowded environments within narrow conduits under external flow conditions, enabling them to perform interesting translational and rotational maneuvers, such as swimming in the upstream…

Soft Condensed Matter · Physics 2026-05-26 Margam Ramprasad , Shubhadeep Mandal , Pallab Sinha Mahapatra

Small objects can swim by generating around them fields or gradients which in turn induce fluid motion past their surface by phoretic surface effects. We quantify for arbitrary swimmer shapes and surface patterns, how efficient swimming…

Soft Condensed Matter · Physics 2009-11-13 R. Golestanian , T. B. Liverpool , A. Ajdari

Living microorganisms are capable of a tactic response to external stimuli by swimming towards or away from the stimulus source; they do so by adapting their tactic signal transduction pathways to the environment. Their self-motility thus…

Soft Condensed Matter · Physics 2016-10-12 Alexander Geiseler , Peter Hänggi , Fabio Marchesoni , Colm Mulhern , Sergey Savel'ev

Microswimmers are sub-millimeter swimming microrobots that show potential as a platform for controllable locomotion in applications including targeted cargo delivery and minimally invasive surgery. To be viable for these target…

Robotics · Computer Science 2024-12-20 Taryn Imamura , Teresa A. Kent , Rebecca E. Taylor , Sarah Bergbreiter

Biological microswimmers often encounter deformable boundaries in physiological conditions; for instance, the viscoelastic walls of reproductive tract during migration of spermatozoa, or host tissue during early bacterial biofilm formation.…

Soft Condensed Matter · Physics 2025-08-07 Smita S. Sontakke , Aneesha Kajampady , Mohd Suhail Rizvi , Ranabir Dey

In this review, we provide a theoretical introduction to Jeffery's equations for the orientation dynamics of an axisymmetric object in a flow at low Reynolds number, and review recent theoretical extensions and applications to the motions…

Fluid Dynamics · Physics 2023-05-31 Kenta Ishimoto

We study different types of microswimmers moving in channels with varying cross section and thereby interacting hydrodynamically with the channel walls. Starting from the Smoluchowski equation for a dilute suspension, for which interactions…

Soft Condensed Matter · Physics 2017-05-24 Paolo Malgaretti , Holger Stark

Microorganisms are rarely found in Nature swimming freely in an unbounded fluid. Instead, they typically encounter other organisms, hard walls, or deformable boundaries such as free interfaces or membranes. Hydrodynamic interactions between…

Fluid Dynamics · Physics 2013-10-21 Marcelo A. Dias , Thomas R. Powers