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Interfaces between stratified epithelia and their supporting stromas commonly exhibit irregular shapes. Undulations are particularly pronounced in dysplastic tissues and typically evolve into long, finger-like protrusions in carcinomas. In…

Biological Physics · Physics 2013-06-25 Thomas Risler , Markus Basan

Multicellular tumor spheroids are an important {\it in vitro} model of the pre-vascular phase of solid tumors, for sizes well below the diagnostic limit: therefore a biophysical model of spheroids has the ability to shed light on the…

Tissues and Organs · Quantitative Biology 2015-05-20 Edoardo Milotti , Roberto Chignola

The internal and external mechanical environment plays an important role in tumorogenesis. As a proxy of an avascular early state tumor, we use multicellular spheroids, a composite material made of cells, extracellular matrix and permeating…

Morphological instabilities of growing tissues that impinge on passive materials are typical of invasive cancers. To explain these instabilities in experiments on breast epithelial spheroids in an extracellular matrix, we develop a…

Soft Condensed Matter · Physics 2026-02-12 Sumit Sinha , Haiqian Yang , L Mahadevan

Spheroids are aggregates of cells that can mimic the cellular organization often found in tissues. They are typically formed through the self-assembly of cells in a culture where there is a promotion of interactions and cell-to-cell…

Signal Processing · Electrical Eng. & Systems 2024-05-24 Ibrahim Isik , Mitra Rezaei , Adam Noel

A simple one-dimensional mechanical model is proposed for splitting instability in swollen membranes. The splitting instability occurs by ring constriction. The bifurcation can be both subcritical and supercritical, depending on the…

Soft Condensed Matter · Physics 2015-06-18 Hidetsugu Sakaguchi , Satomi Maeyama

One of the hallmarks of pre-migratory tumors is the progressive loss of compact morphology. To investigate how tumors may intrinsically regulate their shape during growth, we employ a three-dimensional (3D) vertex model of multicellular…

Soft Condensed Matter · Physics 2026-02-09 Urban Železnik , Matej Krajnc , Tanmoy Sarkar

Tissue surface tension influences cell sorting and tissue fusion. Earlier mechanical studies suggest that multicellular spheroids actively reinforce their surface tension with applied force. Here we study this open question through…

We extend the continuum theory of active nematic fluids to study cell flows and tissue dynamics inside multicellular spheroids, spherical, self-assembled aggregates of cells that are widely used as model systems to study tumour dynamics.…

Soft Condensed Matter · Physics 2022-08-18 Liam J Ruske , Julia M Yeomans

The multicellular organization of diverse systems, such as embryos, intestines and tumours, relies on the coordinated migration of cells in 3D curved environments. In these settings, cells establish supracellular patterns of motion,…

Biological Physics · Physics 2021-10-28 Tom Brandstätter , David B. Brückner , Yu Long Han , Ricard Alert , Ming Guo , Chase P. Broedersz

The collective chemotaxis of multicellular clusters is an important phenomenon in various physiological contexts, ranging from embryonic development to cancer metastasis. Such clusters often display interesting shape dynamics and…

Tissues and Organs · Quantitative Biology 2025-06-03 Monika Sanoria , Gema Malet-Engra , Giorgio Scita , Nir Gov , Ajay Gopinathan

Three-dimensional cultures of cells are gaining popularity as an in vitro improvement over 2D Petri dishes. In many such experiments, cells have been found to organize in aggregates. We present new results of three-dimensional in vitro…

Cell Behavior · Quantitative Biology 2018-11-01 Federica Bubba , Camille Pouchol , Nathalie Ferrand , Guillaume Vidal , Luis Almeida , Benoıt Perthame , Michèle Sabbah

Clinically, palpation is one of the important diagnostic methods to assess tumor malignancy. In laboratory research, it is well accepted that the bulk stiffness of the tumor and the surrounding tissue is closely correlated with the…

Soft Condensed Matter · Physics 2025-09-23 Mrinal Pandey , Bangguo Zhu , Kaitlyn Roach , Young Joon Suh , Jeffrey E Segall , Chung-Yuen Hui , Mingming Wu

Although collective cell migration (CCM) is a highly coordinated and fine-tuned migratory mode, instabilities in the form of cell swirling motion (CSM) often occur. The CSM represents a product of the active turbulence obtained at low…

Tissues and Organs · Quantitative Biology 2021-11-01 Ivana Pajic-Lijakovic , Milan Milivojevic

Growing experimental evidence highlights the relevant role of mechanics in the physiology of solid tumours, even in their early stages. While most of the mathematical models describe tumour growth as a volumetric increase of mass in the…

Biological Physics · Physics 2025-05-19 D. Riccobelli

We develop a self-consistent theory of temporal fluctuations of a speckle pattern resulting from the multiple scattering of a coherent wave in a weakly nonlinear disordered medium. The speckle pattern is shown to become unstable if the…

Disordered Systems and Neural Networks · Physics 2016-08-31 S. E. Skipetrov , R. Maynard

The complex physics of self-assembly in colloidal crystals on deformable interfaces and surfaces poses interesting possibilities for the designability and synthesis of next-generation metamaterials. The goal of this article is to…

Soft Condensed Matter · Physics 2025-06-23 Sanjay Dharmavaram , Basant Lal Sharma

Cell spheroids are a widely used model to investigate cell-cell and cell-matrix interactions in a 3D microenvironment in vitro. Most research on cell spheroids has been focused on their response to various stimuli under static conditions.…

Cell Behavior · Quantitative Biology 2024-04-12 Rosalia Ferraro , Jasmin Di Franco , Sergio Caserta , Stefano Guido

The rheological properties of biological tissues are core to processes such as cancer metastasis, wound healing and embryo development. The emergence of tissue and organ structures during morphogenesis requires the precise formation of…

Soft Condensed Matter · Physics 2026-03-10 Aidan J. Nicholas , Suzanne M. Fielding

Collective cell movement, characterized by multiple cells that are in contact for substantial periods of time and undergo correlated motion, plays a central role in cancer and embryogenesis. Recent imaging experiments have provided…

Biological Physics · Physics 2020-06-03 Sumit Sinha , Abdul N Malmi-Kakkada , Xin Li , Himadri S. Samanta , D. Thirumalai
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