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The thermodynamic (TD) properties of biological membranes play a central role for living systems. It has been suggested, for instance, that nonlinear pulses such as action potentials (APs) can only exist if the membrane state is in vicinity…

Biological Physics · Physics 2020-07-16 Christian Fillafer , Anne Paeger , Matthias F. Schneider

FoF1-ATP synthases are ubiquitous membrane-bound, rotary motor enzymes that can catalyze ATP synthesis and hydrolysis. Their enzyme kinetics are controlled by internal subunit rotation, by substrate and product concentrations, by mechanical…

Biomolecules · Quantitative Biology 2021-06-29 Thomas Heitkamp , Michael Börsch

FoF1-ATP synthase is the membrane protein catalyzing the synthesis of the 'biological energy currency' adenosine triphosphate (ATP). The enzyme uses internal subunit rotation for the mechanochemical conversion of a proton motive force to…

Biomolecules · Quantitative Biology 2015-06-15 Thomas Heitkamp , Hendrik Sielaff , Anja Korn , Marc Renz , Nawid Zarrabi , Michael Boersch

Studies by microbiologists from the 1970s provided robust estimates for the energy supply and demand of a prokaryotic cell. The amount of ATP needed to support growth was calculated from the chemical composition of the cell and known…

Molecular Networks · Quantitative Biology 2025-03-24 William F. Martin

Cellular metabolism is globally regulated by various currency metabolites such as ATP, GTP, and NAD(P)H. These metabolites cycle between charged (high-energy) and uncharged (low-energy) states to mediate energy transfer. While distinct…

Biological Physics · Physics 2026-05-22 Jumpei F. Yamagishi , Tetsuhiro S. Hatakeyama

Understanding the thermodynamic function of life may shed light on its origin. Life, as are all irreversible processes, is contingent on entropy production. Entropy production is a measure of the rate of the tendency of Nature to explore…

General Physics · Physics 2014-03-03 K. Michaelian

The nucleotides ATP and ADP regulate many aspects of endothelial cell (EC) biology, including intracellular calcium concentrations, focal adhesion activation, cytoskeletal organization, and cellular motility. In vivo, ECs are constantly…

Biological Physics · Physics 2018-09-05 Ezio Di Costanzo , Abdul I. Barakat , Giuseppe Pontrelli

Understanding the thermodynamics of the duplication process is a fundamental step towards a comprehensive physical theory of biological systems. However, the immense complexity of real cells obscures the fundamental tensions between energy…

Statistical Mechanics · Physics 2018-12-07 Bernat Corominas-Murtra

Proteases regulate various aspects of the life cycle in all organisms by cleaving specific peptide bonds. Their action is so central for biochemical processes that at least 2% of any known genome encodes for proteolytic enzymes. Here we…

Biomolecules · Quantitative Biology 2013-06-07 Vincenzo Carnevale , Simone Raugei , Cristian Micheletti , Paolo Carloni

Mismatch repair is a critical step in DNA replication that occurs after base selection and proofreading, significantly increasing fidelity. However, the mechanism of mismatch recognition has not been established for any repair enzyme.…

Biological Physics · Physics 2024-04-09 Nianqin Zhang , Yongjun Zhang

ATP-driven proton pumps, which are critical to the operation of a cell, maintain cytosolic and organellar pH levels within a narrow functional range. These pumps employ two very different mechanisms: an elaborate rotary mechanism used by…

Subcellular Processes · Quantitative Biology 2017-11-01 Ramu Anandakrishnan , Daniel M. Zuckerman

F$_{1}$-ATPase is a rotary molecular motor that \emph{in vivo} is subject to strong nonequilibrium driving forces. There is great interest in understanding the operational principles governing its high efficiency of free-energy…

Statistical Mechanics · Physics 2022-12-29 Deepak Gupta , Steven J. Large , Shoichi Toyabe , David A. Sivak

ATPases cyclically convert chemical energy in the form of ATP gradients into directed motion inside cells. To function, ATPases rely on allosteric communication between at least two binding sites, an internal signaling mechanism that is not…

Molecular Networks · Quantitative Biology 2025-12-22 Tosan Omabegho

Adenosine-5'-triphosphate (ATP) plays a crucial role in many biocatalytic reactions and its regeneration can influence the performance of a related enzymatic reaction significantly. Here, we established an electrochemically coupled ATP…

Biomolecules · Quantitative Biology 2023-07-13 Regine Siedentop , Tobias Prenzel , Siegfried R. Waldvogel , Katrin Rosenthal , Stephan Lütz

FoF1-ATP synthases are membrane-embedded protein machines that catalyze the synthesis of adenosine triphosphate. Using photoactivation-based localization microscopy (PALM) in TIR-illumination as well as structured illumination microscopy…

Biomolecules · Quantitative Biology 2015-06-11 Anja Renz , Marc Renz , Diana Kluetsch , Gabriele Deckers-Hebestreit , Michael Börsch

To sustain metabolism, intracellular ATP concentration must be regulated within an appropriate range. This coordination is achieved through the function of the AMP-activated protein kinase (AMPK), a cellular "fuel gauge" that is expressed…

Tissues and Organs · Quantitative Biology 2019-11-07 Benoit Viollet

Inside every living cell is the cytoplasm: a fluid mixture of thousands of different macromolecules, predominantly proteins. This mixture is where most of the biochemistry occurs that enables living cells to function, and it is perhaps the…

Soft Condensed Matter · Physics 2009-11-11 Richard P. Sear

Fo-ATP synthase (Fo) is a rotary motor that converts potential energy from ions, usually protons, moving from high- to low-potential sides of a membrane into torque and rotary motion. Here we propose a mechanism whereby electric fields…

Biological Physics · Physics 2013-09-13 John H. Miller, , Kimal I. Rajapakshe , Hans L. Infante , James R. Claycomb

Kinesin and related motor proteins utilize ATP fuel to propel themselves along the external surface of microtubules in a processive and directional fashion. We show that the observed step-like motion is possible through time varying charge…

Biomolecules · Quantitative Biology 2007-05-23 A. Ciudad , J. M. Sancho , G. P. Tsironis

Trigger waves are self-regenerating propagating fronts that emerge from the coupling of nonlinear reaction kinetics and diffusion. In cells, trigger waves coordinate large-scale processes such as mitotic entry and stress responses. Although…

Molecular Networks · Quantitative Biology 2026-03-12 Jianwei Li , Kai Meng , Xuewen Shen , Fangting Li