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The paper presents a two-phase hydrodynamic model for the numerical simulation of collective motion in a thin layer of active colloids containing spherical microswimmers. The model accounts for three fundamental mechanisms governing the…

Soft Condensed Matter · Physics 2025-12-10 A. Kiverin , S. Luguev , I. Yakovenko

Complex or hostile environments can sometimes inhibit the movement capabilities of diffusive particles or active swimmers, who may thus become stuck in fixed positions. This occurs, for example, in the adhesion of bacteria to surfaces at…

Statistical Mechanics · Physics 2024-01-12 Luca Angelani

We show that activity and broken fore-aft shape symmetry enable microswimmers to cross streamlines in nonuniform shear, a key yet overlooked factor in active cross-stream migration. Using a model of flagellated microswimmers in microchannel…

Soft Condensed Matter · Physics 2025-04-24 Derek C. Gomes , Tapan C. Adhyapak

The swimming behavior of bacteria and other microorganisms is sensitive to the physical properties of the fluid in which they swim. Mucus, biofilms, and artificial liquid-crystalline solutions are all examples of fluids with some degree of…

Soft Condensed Matter · Physics 2014-12-17 Madison S. Krieger , Saverio E. Spagnolie , Thomas R. Powers

Geometric confinement plays an important role in the dynamics of natural and synthetic microswimmers from bacterial cells to self-propelled particles in high-throughput microfluidic devices. However, little is known about the effects of…

Fluid Dynamics · Physics 2014-11-13 Alan Cheng Hou Tsang , Eva Kanso

We study two microswimmers consisting of a spherical rigid head and a passive elastic tail. In the first one the tail is clamped to the head, and the system oscillates under the action of an external torque. In the second one, head and tail…

Fluid Dynamics · Physics 2017-02-02 Giancarlo Cicconofri , Antonio DeSimone

Microorganisms and synthetic microswimmers often encounter complex environments consisting of networks of obstacles embedded into viscous fluids. Such settings include biological media, such as mucus with filamentous networks, as well as…

Micron-sized self-propelled (active) particles can be considered as model systems for characterizing more complex biological organisms like swimming bacteria or motile cells. We produce asymmetric microswimmers by soft lithography and study…

We experimentally, numerically and analytically explore the diffusive transport of active colloidal particles with sensory delay, navigating motility landscapes in which the self-propulsion speed depends on space. We show how the transport…

Statistical Mechanics · Physics 2026-04-29 Adrià Garcés , Ueli Töpfer , Lucio Isa , Demian Levis , Ignacio Pagonabarraga

The diffusion type is determined not only by microscopic dynamics but also by the environment properties. For example, the environment's fractal structure is responsible for the emergence of subdiffusive scaling of the mean square…

Statistical Mechanics · Physics 2021-10-26 Piotr Kubala , Michał Cieśla , Bartłomiej Dybiec

Catalytic microswimmers convert the chemical energy of a fuel into motion, sustaining spatial chemical gradients and fluid flows that drive their propulsion. This leads to unconventional individual behavior and the emergence of collective…

Soft Condensed Matter · Physics 2024-09-04 Celso Carrasco , Quentin Martinet , Zaiyi Shen , Juho S. Lintuvuori , Jérémie Palacci , Antoine Aubret

In addition to enabling movement towards environments with favourable living conditions, swimming by microorganisms has also been linked to enhanced mixing and improved nutrient uptake by their populations. Experimental studies have shown…

Fluid Dynamics · Physics 2017-11-07 Blaise Delmotte , Eric E Keaveny , Eric Climent , Franck Plouraboué

Biological microswimmers are known to navigate upstream of an external flow (positive rheotaxis) in trajectories ranging from linear, spiral to oscillatory. Such rheotaxis stems from the interplay between the motion and complex shapes of…

Soft Condensed Matter · Physics 2022-05-27 Ranabir Dey , Carola M. Buness , Babak Vajdi Hokmabad , Chenyu Jin , Corinna C. Maass

We study a model of diffusive oscillators whose internal states are subject to a periodic drive. These models are inspired by the dynamics of deformable particles with pulsating sizes, where repulsion leads to arrest the internal pulsation…

Statistical Mechanics · Physics 2025-05-15 Alessandro Manacorda , Étienne Fodor

Motivated by recent experiments, we consider the hydrodynamic capture of a microswimmer near a stationary spherical obstacle. Simulations of model equations show that a swimmer approaching a small spherical colloid is simply scattered. In…

Fluid Dynamics · Physics 2015-05-12 Saverio E. Spagnolie , Gregorio R. Moreno-Flores , Denis Bartolo , Eric Lauga

We illustrate a concept for shape-changing microswimmers, which exploits the hysteresis of a shape transition of an elastic object, by an elastic disk undergoing cyclic localized swelling. Driving the control parameter of a hysteretic shape…

Soft Condensed Matter · Physics 2020-12-29 Christian Wischnewski , Jan Kierfeld

We numerically investigate the motion of active artificial microswimmers diffusing in a fuel concentration gradient. We observe that, in the steady state, their probability density accumulates in the low-concentration regions, whereas a…

Soft Condensed Matter · Physics 2025-08-18 Pulak K. Ghosh , Yunyun Li , Fabio Marchesoni , Franco Nori

Microswimmers play an important role in shaping the world around us. The squirmer is a simple model for microswimmer whose cilia oscillations on its spherical surface induce an effective slip velocity to propel itself. The rapid development…

Fluid Dynamics · Physics 2024-11-22 Xinwei Cai , Kuiliang Wang , Gaojin Li , Xin Bian

In this paper we deal with pedestrian modeling, aiming at simulating crowd behavior in normal and emergency scenarios, including highly congested mass events. We are specifically concerned with a new agent-based, continuous-in-space,…

Adaptation and Self-Organizing Systems · Physics 2023-11-23 E. Cristiani , M. Menci , A. Malagnino , G. G. Amaro

In isotropic fluids like water, micrometer-scale swimmers have evolved swim strokes to translate despite their tiny size. As described by Purcell in his Scallop Theorem, reciprocal motions, like those performed by a scallop, cannot drive…