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F1F0 ATP synthase (ATPase) either facilitates the synthesis of ATP in the mitochondrial membranes and bacterial inner membranes in a process driven by the proton moving force (pmf), or uses the energy from ATP hydrolysis to pump protons…

Subcellular Processes · Quantitative Biology 2016-07-12 O. Kulish , A. D. Wright , E. M. Terentjev

The proton motive force (PMF) across the inner mitochondrial membrane delivers approximately 0.2 eV of energy per proton, powering the FoF1-ATP synthase molecular motor. Here, we provide a detailed accounting of how this energy is utilized:…

Biomolecules · Quantitative Biology 2025-07-16 Islam K. Matar , Peyman Fahimi , Cherif F. Matta

The diffusion of a molecular motor in the presence of a constant external force is considered on the basis of a simple theoretical model. The motor is represented by a Brownian particle moving in a series of parabolic potentials placed…

Statistical Mechanics · Physics 2016-06-01 Ryota Shinagawa , Kazuo Sasaki

Many swimming bacteria are propelled by flagellar motors that stochastically switch between the clockwise and counterclockwise rotation direction. While the switching dynamics are one of the most important characteristics of flagellar…

Subcellular Processes · Quantitative Biology 2008-11-18 Siebe B. van Albada , Sorin Tanase-Nicola , Pieter Rein ten Wolde

The pump process of the ratchet model inspired by the $F_o$ rotatory motor of ATP synthase is investigated. In this model there are two kinds of characteristic time. One is dynamical, the relaxation time of the system. Others are chemical,…

Biological Physics · Physics 2007-05-23 Hiroshi Miki , Masatoshi Sato , Mahito Kohmoto

Molecular motors fulfill critical functions within all living beings. Understanding their underlying working principles is therefore of great interest. Here we develop a simple model inspired by the two-component biomolecular motor Fo-F1…

Statistical Mechanics · Physics 2024-12-30 Mathis Grelier , David A. Sivak , Jannik Ehrich

The motility of certain gram-negative bacteria is mediated by retraction of type IV pili surface filaments, which are essential for infectivity. The retraction is powered by a strong molecular motor protein, PilT, producing very high forces…

Subcellular Processes · Quantitative Biology 2007-05-23 Martin Linden , Tomi Tuohimaa , Ann-Beth Jonsson , Mats Wallin

In this work we study the assisted translocation of a polymer across a membrane nanopore, inside which a molecular motor exerts a force fuelled by the hydrolysis of ATP molecules. In our model the motor switches to its active state for a…

Soft Condensed Matter · Physics 2018-03-26 A. Fiasconaro , J. J. Mazo , F. Falo

The type IV pilus retraction motor is found in many important bacterial pathogens. It is the strongest known linear motor protein and is required for bacterial infectivity. We characterize the dynamics of type IV pilus retraction in terms…

Biological Physics · Physics 2007-05-23 Martin Linden , Emil Johansson , Ann-Beth Jonsson , Mats Wallin

The enzyme FoF1-ATP synthase provides the 'chemical energy currency' adenosine triphosphate (ATP) for living cells. Catalysis is driven by mechanochemical coupling of subunit rotation within the enzyme with conformational changes in the…

Biomolecules · Quantitative Biology 2015-06-04 Stefan Ernst , Monika G. Dueser , Nawid Zarrabi , Michael Boersch

Two simple (rotator and one-particle) mechanistic models are suggested to describe simultaneously at a minimal level of sophistication two basic functions of F$_1$-ATPase: a motor regime driven by ATP hydrolysis and its inverted function as…

Biological Physics · Physics 2007-05-23 A. V. Zolotaryuk , V. N. Ermakov , P. L. Christiansen , B. Norden , Y. Zolotaryuk

FoF1-ATP synthases are ubiquitous membrane-bound, rotary motor enzymes that can catalyze ATP synthesis and hydrolysis. Their enzyme kinetics are controlled by internal subunit rotation, by substrate and product concentrations, by mechanical…

Biomolecules · Quantitative Biology 2021-06-29 Thomas Heitkamp , Michael Börsch

F$_\mathrm{o}$F$_1$-ATP synthase is a factory for synthesizing ATP in virtually all cells. Its core machinery is the subcomplex F$_1$-motor (F$_1$-ATPase) and performs the reversible mechanochemical coupling. Isolated F$_1$-motor hydrolyzes…

Biological Physics · Physics 2015-01-19 Shoichi Toyabe , Eiro Muneyuki

We demonstrate asymmetric enzyme kinetics of a biomolecular motor F1-ATPase between synthesis and hydrolysis of adenosine triphosphate (ATP). Our experiments show that ATP hydrolysis follows Michaelis-Menten kinetics, but ATP synthesis,…

Biological Physics · Physics 2025-06-04 Yohei Nakayama , Shoichi Toyabe

F$_1$-ATPase (F$_1$) is central to cellular energy transduction. Forcibly rotated by another motor F$_\mathrm{o}$, F$_1$ catalyzes ATP synthesis by converting mechanical work into chemical free energy stored in the molecule ATP. The details…

FoF1-ATP synthase is the enzyme that provides the 'chemical energy currency' adenosine triphosphate, ATP, for living cells. The formation of ATP is accomplished by a stepwise internal rotation of subunits within the enzyme. Briefly, proton…

Biomolecules · Quantitative Biology 2009-11-13 N. Zarrabi , S. Ernst , M. G. Dueser , A. Golovina-Leiker , W. Becker , R. Erdmann , S. D. Dunn , M. Borsch

The bacterial flagellar motor (BFM) converts transmembrane ion flux into directed mechanical rotation, driving bacterial motility. Despite extensive study, the frictional forces and energetics governing its torque generation remain poorly…

Biological Physics · Physics 2025-12-18 N. J. Lopez-Alamilla , A. L. Nord , F. Pedaci , J. Palmeri , N. -O. Walliser

We have used numerical simulations to investigate how the properties of motor proteins control the dynamical behavior of a driven flexible filament. The filament is pinned at one end and positioned on top of a patch of anchored motor…

Soft Condensed Matter · Physics 2024-09-20 Amir Khosravanizadeh , Serge Dmitrieff

Molecular motors drive mechanical motions utilizing the free energy liberated from chemical reactions such as ATP hydrolysis. Although it is essential to know the efficiency of this free energy transduction, it has been a challenge due to…

A discrete-state model of the F1-ATPase molecular motor is developed which describes not only the dependences of the rotation and ATP consumption rates on the chemical concentrations of ATP, ADP, and inorganic phosphate, but also on…

Subcellular Processes · Quantitative Biology 2009-04-28 E. Gerritsma , P. Gaspard
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