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Recent work on stochastic interacting particle systems with two particle species (or single-species systems with kinematic constraints) has demonstrated the existence of spontaneous symmetry breaking, long-range order and phase coexistence…

Statistical Mechanics · Physics 2016-08-16 Gunter M. Schütz

Despite their importance in many biological, ecological and physical processes, microorganismal fluid flows under tight confinement have not been investigated experimentally. Strong screening of Stokelets in this geometry suggests that the…

Biological Physics · Physics 2019-12-18 Raphaël Jeanneret , Dmitri O. Pushkin , Marco Polin

Many aquatic microorganisms are able to swim. In natural environments they typically do so in the presence of flows. In recent years it has been shown that the interplay of swimming and flows can give rise to interesting and biologically…

Fluid Dynamics · Physics 2022-07-20 M. Borgnino , G. Boffetta , M. Cencini , F. De Lillo , K. Gustavsson

Microtubules and motor proteins are building blocks of self-organized subcellular biological structures such as the mitotic spindle and the centrosomal microtubule array. These same ingredients can form new "bioactive" liquid-crystalline…

Soft Condensed Matter · Physics 2015-01-29 Tong Gao , Robert Blackwell , Matthew A. Glaser , M. D. Betterton , Michael J. Shelley

Organisms often swim through density stratified fluids. In this Letter, we investigate the dynamics of small active particles swimming in density gradients and report theoretical evidence of taxis as a result of density stratification…

Soft Condensed Matter · Physics 2023-08-02 Vaseem A. Shaik , Gwynn J. Elfring

Many active matter systems are known to perform L\'{e}vy walks during migration or foraging. Such superdiffusive transport indicates long-range correlated dynamics. These behavior patterns have been observed for microswimmers such as…

Soft Condensed Matter · Physics 2021-05-28 Tingtao Zhou , Zhiwei Peng , Mamikon Gulian , John F. Brady

Active particles, which interact hydrodynamically, display a remarkable variety of emergent collective phenomena. We use squirmers to model spherical microswimmers and explore the collective behavior of thousands of them under the influence…

Fluid Dynamics · Physics 2017-10-31 Jan-Timm Kuhr , Johannes Blaschke , Felix Rühle , Holger Stark

We develop a dynamical density functional theory based model for the drying of colloidal films on planar surfaces. We consider mixtures of two different sizes of hard-sphere colloids. Depending on the solvent evaporation rate and the…

Soft Condensed Matter · Physics 2021-01-28 Boshen He , Ignacio Martin-Fabiani , Roland Roth , Gyula I. Tóth , Andrew J. Archer

Guiding active microswimmers by external fields to requested target locations is a promising strategy to realize complex transport on the microscale. To this end, one possibility consists of attaching the microswimmers to orientable passive…

Fluid Dynamics · Physics 2020-02-19 Abdallah Daddi-Moussa-Ider , Maciej Lisicki , Hartmut Löwen , Andreas M. Menzel

Biological microswimmers alter their motility in complex corner geometries, facilitating their survival. However, the dynamical features of low-Reynolds-number swimming at corners remain undefined. Here, we use active droplet microswimmers…

Soft Condensed Matter · Physics 2026-04-22 Subhasish Guchhait , Harshita Tiwari , Sumesh P. Thampi , Ranabir Dey

One approach to quantifying biological diversity consists of characterizing the statistical distribution of specific properties of a taxonomic group or habitat. Microorganisms living in fluid environments, and for whom motility is key,…

Soft Condensed Matter · Physics 2020-12-03 Maciej Lisicki , Marcos F. Velho Rodrigues , Raymond E. Goldstein , Eric Lauga

Chiral active particles, or self-propelled circle swimmers, from sperm cells to asymmetric Janus colloids, form a rich set of patterns, which are different from those seen in linear swimmers. Such patterns have mainly been explored for…

Soft Condensed Matter · Physics 2019-07-24 Demian Levis , Benno Liebchen

Hydrodynamic interactions driven by particle activity are ubiquitous in active colloidal systems. Although these interactions are strongly influenced by the interfacial actuation mechanism and geometry of the swimming particles, theoretical…

Soft Condensed Matter · Physics 2025-06-17 Anson G. Thambi , William E. Uspal

Many types of mammalian cells exert active contractile forces and mechanically deform their elastic substrate, to accomplish biological functions such as cell migration. These substrate deformations provide a mechanism by which cells can…

Soft Condensed Matter · Physics 2022-02-22 Subhaya Bose , Patrick S. Noerr , Ajay Gopinathan , Arvind Gopinath , Kinjal Dasbiswas

We study two microswimmers consisting of a spherical rigid head and a passive elastic tail. In the first one the tail is clamped to the head, and the system oscillates under the action of an external torque. In the second one, head and tail…

Fluid Dynamics · Physics 2017-02-02 Giancarlo Cicconofri , Antonio DeSimone

We show that a two-dimensional system of flocking microswimmers interacting hydrodynamically can be expressed using a Hamiltonian formalism. The Hamiltonian depends strictly on the angles between the particles and their swimming…

Soft Condensed Matter · Physics 2023-05-23 Yuval Shoham , Naomi Oppenheimer

The energy dissipation and entropy production by self-propelled microswimmers differ profoundly from passive particles pulled by external forces. The difference extends both to the shape of the flow around the swimmer, as well as to the…

Soft Condensed Matter · Physics 2023-10-02 Abdallah Daddi-Moussa-Ider , Ramin Golestanian , Andrej Vilfan

A universal characteristic of mesoscale turbulence in active suspensions is the emergence of a typical vortex length scale, distinctly different from the scale-invariance of turbulent high-Reynolds number flows. Collective length-scale…

Soft Condensed Matter · Physics 2016-08-17 Sebastian Heidenreich , Jörn Dunkel , Sabine H. L. Klapp , Markus Bär

Classical swarm models, exemplified by the Cucker--Smale framework, provide foundational insights into collective alignment but exhibit fundamental limitations in capturing the adaptive, heterogeneous behaviours intrinsic to living systems.…

Adaptation and Self-Organizing Systems · Physics 2025-09-08 Rene Fabregas , Jie Liao , Nisrine Outada

Understanding the transport properties of microorganisms and self-propelled particles in porous media has important implications for human health as well as microbial ecology. In free space, most microswimmers perform diffusive random walks…

Soft Condensed Matter · Physics 2023-08-10 David Saintillan