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In equilibrium, the collective behaviour of particles interacting via steep, short-ranged potentials is well captured by the virial expansion of the free energy at low density. Here, we extend this approach beyond equilibrium to the case of…

Soft Condensed Matter · Physics 2023-06-21 Yuting Irene Li , Rosalba Garcia-Millan , Michael E. Cates , Étienne Fodor

Self-propelled active particles exhibit delayed responses to environmental changes, modulating their propulsion speed through intrinsic sensing and feedback mechanisms. This adaptive behavior fundamentally determines their dynamics and…

Soft Condensed Matter · Physics 2025-05-21 Viktor Holubec , Alexander Fischer , Giovanni Volpe , Frank Cichos

In addition to enabling movement towards environments with favourable living conditions, swimming by microorganisms has also been linked to enhanced mixing and improved nutrient uptake by their populations. Experimental studies have shown…

Fluid Dynamics · Physics 2017-11-07 Blaise Delmotte , Eric E Keaveny , Eric Climent , Franck Plouraboué

Shear-thinning is an important rheological property of many biological fluids, such as mucus, whereby the apparent viscosity of the fluid decreases with shear. Certain microscopic swimmers have been shown to progress more rapidly through…

Fluid Dynamics · Physics 2013-09-06 Thomas D. Montenegro-Johnson , Daniel Loghin , David J. Smith

Consider a fluid composed of two species of particles, where the interparticle pair potentials $u_{11} = u_{22} \neq u_{12}$. On confining an equal number of particles from each species in a cavity, one finds that the average one body…

Soft Condensed Matter · Physics 2007-05-23 A. J. Archer

The diffusion of active microscopic organisms in complex environments plays an important role in a wide range of biological phenomena from cell colony growth to single organism transport. Here, we investigate theoretically and…

Fluid Dynamics · Physics 2018-01-16 Juan L. Aragones , Shahrzad Yazdi , Alfredo Alexander-Katz

The effect of gravity on the collective motion of living microswimmers, such as bacteria and micro-algae, is pivotal to unravel not only bio-convection patterns but also the settling of bacterial biofilms on solid surfaces. In this work, we…

Soft Condensed Matter · Physics 2025-04-07 C. Miguel Barriuso G. , Horacio Serna , Ignacio Pagonabarraga , Chantal Valeriani

Turbulence is ubiquitous, from oceanic currents to small-scale biological and quantum systems. Self-sustained turbulent motion in microbial suspensions presents an intriguing example of collective dynamical behavior amongst the simplest…

We study the trajectories of a model microorganism inside three-dimensional channels with square and rectangular cross-sections. Using (i) numerical simulations based on lattice-Boltzmann method, and (ii) analytical expressions using…

Soft Condensed Matter · Physics 2024-08-16 Byjesh N. Radhakrishnan , Ahana Purushothaman , Ranabir Dey , Sumesh P Thampi

Catalytic microswimmers convert the chemical energy of a fuel into motion, sustaining spatial chemical gradients and fluid flows that drive their propulsion. This leads to unconventional individual behavior and the emergence of collective…

Soft Condensed Matter · Physics 2024-09-04 Celso Carrasco , Quentin Martinet , Zaiyi Shen , Juho S. Lintuvuori , Jérémie Palacci , Antoine Aubret

Disordered hyperuniformity, a brand new type of arrangements with novel physical properties, provides various practical applications in extensive fields. To highlight the great potential of applying disordered hyperuniformity to active…

Soft Condensed Matter · Physics 2019-05-30 Jie Su , Huijun Jiang , Zhonghuai Hou

The dynamical properties of classical fluids at pico-liter scale attract experimentally and theoretically much attention in the soft-matter and biophysics communities, due to the appearance of the microfluidics, also called 'lab-on-a-chip',…

Statistical Mechanics · Physics 2007-05-23 F. Penna , P. Tarazona

Biological microswimmers often encounter deformable boundaries in physiological conditions; for instance, the viscoelastic walls of reproductive tract during migration of spermatozoa, or host tissue during early bacterial biofilm formation.…

Soft Condensed Matter · Physics 2025-08-07 Smita S. Sontakke , Aneesha Kajampady , Mohd Suhail Rizvi , Ranabir Dey

We propose a combined analytical-numerical strategy to predict the dynamics and trajectory of a microswimmer next to a curved spherical obstacle. The microswimmer is actuated by a slip velocity on its surface and a uniformly valid solution…

Soft Condensed Matter · Physics 2017-10-31 Nima Sharifi-Mood , Pablo G. Díaz-Hyland , Ubaldo M. Córdova-Figueroa

Low Reynolds number direct simulations of large populations of hydrodynamically interacting swimming particles confined between planar walls are performed. The results of simulations are compared with a theory that describes dilute…

Fluid Dynamics · Physics 2015-05-13 Juan P. Hernandez-Ortiz , Patrick T. Underhill , Michael D. Graham

We theoretically describe the dynamics of swimmer populations confined in thin liquid films. We first demonstrate that hydrodynamic interactions between confined swimmers only depend on their shape and are independent of their specific…

Soft Condensed Matter · Physics 2015-06-11 Tommaso Brotto , Jean-Baptiste Caussin , Eric Lauga , Denis Bartolo

We investigate how shear influences the dynamics of active particles in dilute to concentrated suspensions. Using apolar active suspensions of squirmers as model systems, we show how their long-time diffusive dynamics can surprisingly slow…

Soft Condensed Matter · Physics 2025-11-14 Zhouyang Ge , John F. Brady , Gwynn J. Elfring

We study numerically the hydrodynamics of a self-propelled particle system, consisting of spherical squirmers sedimented on a flat surface. We observe the emergence of dynamic structures, due to the interplay of particle-particle and…

Soft Condensed Matter · Physics 2019-12-20 Zaiyi Shen , Juho S. Lintuvuori

We investigate the hydrodynamic interactions between microorganisms swimming at low Reynolds number. By considering simple model swimmers, and combining analytic and numerical approaches, we investigate the time-averaged flow field around a…

Soft Condensed Matter · Physics 2007-05-25 C. M. Pooley , G. P. Alexander , J. M. Yeomans

We propose minimal models of one-, two- and three-dimensional micro-swimmers at low Reynolds number with a periodic non-reciprocal motion. These swimmers are either "pushers" or "pullers" of fluid along the swimming axis, or combination of…

Soft Condensed Matter · Physics 2010-04-30 Nobuhiko Watari , Ronald G. Larson