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We use numerical simulations to probe the dynamics of concentrated suspensions of spherical microswimmers interacting hydrodynamically. Previous work in the dilute limit predicted orientational instabilities of aligned suspensions for both…

Soft Condensed Matter · Physics 2013-02-12 Arthur A. Evans , Takuji Ishikawa , Takami Yamaguchi , Eric Lauga

Microswimmers typically operate in complex environments. In biological systems, often diverse species are simultaneously present and interact with each other. Here, we derive a (time-dependent) particle-scale statistical description, namely…

Soft Condensed Matter · Physics 2019-08-13 Christian Hoell , Hartmut Löwen , Andreas M. Menzel

Microswimmers are active particles of microscopic size that self-propel by setting the surrounding fluid into motion. According to the kind of far-field fluid flow that they induce, they are classified into pushers and pullers. Many studies…

Soft Condensed Matter · Physics 2019-08-01 Giorgio Pessot , Hartmut Löwen , Andreas M. Menzel

We explore the mechanism underlying the anomalous rheology of puller-type microswimmer suspensions through direct hydrodynamic simulations. Puller-type swimmers generate contractile flow fields along their swimming direction, leading to…

Soft Condensed Matter · Physics 2025-03-04 Haruki Hayano , Akira Furukawa

We investigate the hydrodynamic stability and transport of magnetic microswimmers in an external field using a kinetic theory framework. Combining linear stability analysis and nonlinear 3D continuum simulations, we show that for…

Soft Condensed Matter · Physics 2019-02-20 Fabian R. Koessel , Sara Jabbari-Farouji

We present a comprehensive theory of the dynamics and fluctuations of a two-dimensional suspension of polar active particles in an incompressible fluid confined to a substrate. We show that, depending on the sign of a single parameter, a…

Soft Condensed Matter · Physics 2020-01-22 Ananyo Maitra , Pragya Srivastava , M. Cristina Marchetti , Sriram Ramaswamy , Martin Lenz

In this paper we systematically derive a fourth-order continuum theory capable of reproducing mesoscale turbulence in a three-dimensional suspension of microswimmers. We start from overdamped Langevin equations for a generic microscopic…

Soft Condensed Matter · Physics 2018-06-28 Henning Reinken , Sabine H. L. Klapp , Markus Bär , Sebastian Heidenreich

Self-propelled particles with hydrodynamic interactions (microswimmers) have previously been shown to produce long-range ordering phenomena. Many theoretical explanations for these collective phenomena are connected to instabilities in the…

Soft Condensed Matter · Physics 2017-05-03 Yuzhou Qian , Peter R. Kramer , Patrick T. Underhill

We study numerically the hydrodynamics of a self-propelled particle system, consisting of spherical squirmers sedimented on a flat surface. We observe the emergence of dynamic structures, due to the interplay of particle-particle and…

Soft Condensed Matter · Physics 2019-12-20 Zaiyi Shen , Juho S. Lintuvuori

Hydrodynamical interactions of active micro-particles are pervasive in our planet's fluid environments. Hence, understanding the interactions of these self-propelled particles is essential for science and engineering. In this paper the…

Soft Condensed Matter · Physics 2022-12-28 Yonatan Ashenafi

Systems of motile microorganisms exhibit a multitude of collective phenomena, including motility-induced phase separation and turbulence. Sensing of the environment and adaptation of movement plays an essential role in the emergent…

Soft Condensed Matter · Physics 2024-06-26 Segun Goh , Elmar Westphal , Roland G. Winkler , Gerhard Gompper

Active suspensions encompass a wide range of complex fluids containing microscale energy-injecting particles, such as cells, bacteria or artificially powered active colloids. Because they are intrinsically non-equilibrium, active…

Soft Condensed Matter · Physics 2024-07-09 Zhouyang Ge , Gwynn J. Elfring

Microorganisms living in microfluidic environments often form multi-species swarms, where they can leverage collective motions to achieve enhanced transport and spreading. Nevertheless, there is a general lack of physical understandings of…

Fluid Dynamics · Physics 2022-02-14 Bryce Palmer , Wen Yan , Tong Gao

We identify the presence of a continuum percolation transition in model suspensions of pusher-type microswimmers. The clusters dynamically aggregate and disaggregate resulting from a competition of attractive and repulsive hydrodynamic and…

Soft Condensed Matter · Physics 2019-08-29 Fabian Jan Schwarzendahl , Marco G. Mazza

Microswimmers often exhibit surprising patterns due to the nonequilibrium nature of their dynamics. Collectively, suspensions of microswimmers appear as a liquid whose properties set it apart from its passive counterpart. To understand the…

Soft Condensed Matter · Physics 2019-05-22 Fabian Jan Schwarzendahl , Marco G. Mazza

We construct the hydrodynamic equations for {\em suspensions} of self-propelled particles (SPPs) with spontaneous orientational order, and make a number of striking, testable predictions:(i) SPP suspensions with the symmetry of a true {\em…

Soft Condensed Matter · Physics 2009-11-07 R. Aditi Simha , Sriram Ramaswamy

A collection of microswimmers immersed in an incompressible fluid is characterised by strong interactions due to the long-range nature of the hydrodynamic fields generated by individual organisms. As a result, suspensions of rear-actuated…

Soft Condensed Matter · Physics 2023-12-19 Viktor Škultéty , Dóra Bárdfalvy , Joakim Stenhammar , Cesare Nardini , Alexander Morozov

Microorganisms ofter move in confined, disordered environments, where hydrodynamic couplings can modify their transport behavior. Using extensive finite-element simulations, we investigate the dynamics of microswimmers -- modeled as…

Soft Condensed Matter · Physics 2026-03-24 Mirko Residori , Sebastian Aland , Christina Kurzthaler

The emergence of hydrodynamic bend instabilities in ordered suspensions of active particles is widely observed across diverse living and synthetic systems, and is considered to be governed by dipolar active stresses generated by the…

Soft Condensed Matter · Physics 2026-01-09 Sameer Kumar , Niels de Graaf Sousa , Amin Doostmohammadi

The habitat of microorganisms is typically complex and viscoelastic. A natural question arises: Do polymers in a suspension of active swimmers enhance chaotic motion or promote orientational order? We address this issue by performing…

Soft Condensed Matter · Physics 2025-12-23 Yuan Zhou , Qingzhi Zou , Ignacio Pagonabarraga , Kaihuan Zhang , Kai Qi
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