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This paper studies the influence of orienting external fields on pattern formation, particularly mesoscale turbulence, in microswimmer suspensions. To this end, we apply a hydrodynamic theory that can be derived from a microscopic…

Soft Condensed Matter · Physics 2019-02-27 Henning Reinken , Sebastian Heidenreich , Markus Bär , Sabine H. L. Klapp

Meso-scale turbulence was originally observed experimentally in various suspensions of swimming bacteria, as well as in the collective motion of active colloids. The corresponding large-scale dynamical patterns were reproduced in a simple…

Soft Condensed Matter · Physics 2021-03-30 Vasco M. Worlitzer , Gil Ariel , Avraham Be'er , Holger Stark , Markus Bär , Sebastian Heidenreich

We study how hydrodynamic interactions affect the collective behaviour of active particles suspended in a fluid at high concentrations, with particular attention to lubrication forces which appear when the particles are very close to one…

Soft Condensed Matter · Physics 2018-05-29 Natsuhiko Yoshinaga , Tanniemola B. Liverpool

We study a planar motion of two hydrodynamically coupled non-identical micro-swimmers, each modelled as a force dipole with intrinsic self-propulsion. Using the method of images, we demonstrate that our results remain equally applicable at…

Chaotic Dynamics · Physics 2022-09-14 Prajitha Mottammal , Sumesh P. Thampi , Andrey Pototsky

Microswimmers in suspension exhibit collective swimming behaviour, forming various self-organised structures including ordered, aggregated, and turbulent-like structures. When mixed with passive particles phase-separation is known to occur,…

Fluid Dynamics · Physics 2026-03-26 Alexander Chamolly , Takuji Ishikawa

The hydrodynamic interactions of a suspension of self-propelled particles are studied using a direct numerical simulation method which simultaneously solves for the host fluid and the swimming particles. A modified version of the "Smoothed…

Soft Condensed Matter · Physics 2013-01-15 John J. Molina , Yasuya Nakayama , Ryoichi Yamamoto

Self-propelled microparticles create flow fields that determine how they interact with surfaces, external flows, and each other. These flow fields fall into distinct classes--pushers, pullers, and neutral swimmers--each exhibiting…

Soft Condensed Matter · Physics 2025-12-25 Lisa Rohde , Gordei Anchutkin , Viktor Holubec , Frank Cichos

An artificial microswimmer drifts in response to spatio-temporal modulations of an activating suspension medium. We consider two competing mechanisms capable of influencing its tactic response: angular fluctuations, which help it explore…

Soft Condensed Matter · Physics 2017-02-15 Alexander Geiseler , Peter Hänggi , Fabio Marchesoni

Active matter exhibits various forms of non-equilibrium states in the absence of external forcing, including macroscopic steady-state currents. Such states are often too complex to be modelled from first principles and our understanding of…

Soft Condensed Matter · Physics 2020-09-17 Viktor Škultéty , Cesare Nardini , Joakim Stenhammar , Davide Marenduzzo , Alexander Morozov

Active matter has been the object of huge amount of research in recent years for its important fundamental and applicative properties. In this paper we investigate active suspensions of micro-swimmers through direct numerical simulation, so…

Soft Condensed Matter · Physics 2021-02-03 Sergio Chibbaro , Astrid Decoene , Sebastien Martin , Fabien Vergnet

Deformable boundaries are omnipresent in the habitats of swimming microorganisms, leading to intricate hydroelastic couplings. Employing a perturbation theory, valid for small deformations, we study the swimming dynamics of pushers and…

Soft Condensed Matter · Physics 2025-02-05 Sagnik Garai , Ursy Makanga , Akhil Varma , Christina Kurzthaler

A universal characteristic of mesoscale turbulence in active suspensions is the emergence of a typical vortex length scale, distinctly different from the scale-invariance of turbulent high-Reynolds number flows. Collective length-scale…

Soft Condensed Matter · Physics 2016-08-17 Sebastian Heidenreich , Jörn Dunkel , Sabine H. L. Klapp , Markus Bär

We study low Reynolds number turbulence in a suspension of polar, extensile, self-propelled inertial swimmers. We review the bend and splay mechanisms that destabilize an ordered flock. The suspension is always unstable to bend…

Soft Condensed Matter · Physics 2025-11-10 Purnima Jain , Navdeep Rana , Roberto Benzi , Prasad Perlekar

Populations of swimming microorganisms produce fluid motions that lead to dramatically enhanced diffusion of tracer particles. Using simulations of suspensions of swimming particles in a periodic domain, we capture this effect and show that…

Soft Condensed Matter · Physics 2008-07-22 Patrick T. Underhill , Juan P. Hernández-Ortiz , Michael D. Graham

Hydrodynamic interactions are crucial for determining the cooperative behavior of microswimmers at low Reynolds numbers. Here we provide a comprehensive analysis of the scaling and strength of the interactions in the case of a pair of…

Soft Condensed Matter · Physics 2021-02-11 Sebastian Ziegler , Thomas Scheel , Maxime Hubert , Jens Harting , Ana-Sunčana Smith

We investigate theoretically the collective dynamics of a suspension of low Reynolds number swimmers that are confined to two dimensions by a thin fluid film. Our model swimmer is characterized by internal degrees of freedom which locally…

Biological Physics · Physics 2015-05-19 M. Leoni , T. B. Liverpool

We study the effects of inertia in dense suspensions of polar swimmers. The hydrodynamic velocity field and the polar order parameter field describe the dynamics of the suspension. We show that a dimensionless parameter $R$ (ratio of the…

Soft Condensed Matter · Physics 2024-08-14 Navdeep Rana , Rayan Chatterjee , Sunghan Ro , Dov Levine , Sriram Ramaswamy , Prasad Perlekar

We propose a hydrodynamic theory to examine the emergence of contraction waves in dense active liquids composed of pulsating deformable particles. Our theory couples the liquid density with a chemical phase that determines the periodic…

Statistical Mechanics · Physics 2025-09-24 Tirthankar Banerjee , Thibault Desaleux , Jonas Ranft , Étienne Fodor

We consider a suspension of active rigid particles (swimmers) in a steady Stokes flow, where particles are distributed according to a stationary ergodic random process, and we study its homogenization in the macroscopic limit. A key point…

Analysis of PDEs · Mathematics 2023-03-15 Armand Bernou , Mitia Duerinckx , Antoine Gloria

The experiments of Leptos et al. [Phys. Rev. Lett. 103, 198103 (2009)] show that the displacements of small particles affected by swimming microorganisms achieve a non-Gaussian distribution, which nevertheless scales diffusively -- the…

Soft Condensed Matter · Physics 2017-10-26 Jean-Luc Thiffeault